position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2025/12/18/peak-motifs.2025-12-18.171557_2025-12-18.171557_4Sppd9/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title HG_0.01 -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2025/12/18/peak-motifs.2025-12-18.171557_2025-12-18.171557_4Sppd9/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2025/12/18/peak-motifs.2025-12-18.171557_2025-12-18.171557_4Sppd9/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2025/12/18/peak-motifs.2025-12-18.171557_2025-12-18.171557_4Sppd9/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 2495 Sum of sequence lengths 644090 Min sequence length 0 Max sequence length 1000 Average sequence length 258 Possible positions 641595 Oligonucleotide statistics: Total occurrences 539979 Position interval parameters: Position interval 20 Number of windows 50 Total positions 641595 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 6 120 2 -24 [-479,-460] -469.5 7 132 3 -23 [-459,-440] -449.5 10 178 4 -22 [-439,-420] -429.5 12 224 5 -21 [-419,-400] -409.5 16 286 6 -20 [-399,-380] -389.5 20 364 7 -19 [-379,-360] -369.5 22 404 8 -18 [-359,-340] -349.5 28 492 9 -17 [-339,-320] -329.5 38 670 10 -16 [-319,-300] -309.5 51 899 11 -15 [-299,-280] -289.5 64 1183 12 -14 [-279,-260] -269.5 84 1454 13 -13 [-259,-240] -249.5 112 2008 14 -12 [-239,-220] -229.5 149 2609 15 -11 [-219,-200] -209.5 206 3572 16 -10 [-199,-180] -189.5 303 5077 17 -9 [-179,-160] -169.5 436 7459 18 -8 [-159,-140] -149.5 656 10921 19 -7 [-139,-120] -129.5 1005 16527 20 -6 [-119,-100] -109.5 1634 26346 21 -5 [-99,-80] -89.5 2495 42780 22 -4 [-79,-60] -69.5 2495 49900 23 -3 [-59,-40] -49.5 2495 49900 24 -2 [-39,-20] -29.5 2495 49900 25 -1 [-19,0] -9.5 2495 49900 26 0 [1,20] 10.5 2495 49900 27 1 [21,40] 30.5 2495 49900 28 2 [41,60] 50.5 2495 49900 29 3 [61,80] 70.5 2495 49900 30 4 [81,100] 90.5 2495 41008 31 5 [101,120] 110.5 1575 25092 32 6 [121,140] 130.5 972 15828 33 7 [141,160] 150.5 627 10495 34 8 [161,180] 170.5 424 7187 35 9 [181,200] 190.5 296 4877 36 10 [201,220] 210.5 200 3456 37 11 [221,240] 230.5 145 2538 38 12 [241,260] 250.5 110 1958 39 13 [261,280] 270.5 82 1414 40 14 [281,300] 290.5 63 1162 41 15 [301,320] 310.5 50 870 42 16 [321,340] 330.5 38 652 43 17 [341,360] 350.5 28 481 44 18 [361,380] 370.5 22 402 45 19 [381,400] 390.5 20 355 46 20 [401,420] 410.5 16 277 47 21 [421,440] 430.5 12 222 48 22 [441,460] 450.5 10 172 49 23 [461,480] 470.5 7 130 50 24 [481,500] 490.5 6 114 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| tt | tt | 76444 | 1325.5 | 49 | 0.0e+00 | 0 | 300 | 1 | 2 | 2 | 4 | 12 | 9 | 11 | 14 | 19 | 31 | 49 | 68 | 88 | 143 | 210 | 276 | 465 | 792 | 1082 | 1825 | 2918 | 4803 | 6310 | 6285 | 6403 | 6101 | 6260 | 6442 | 6550 | 6522 | 5005 | 2934 | 1687 | 1105 | 719 | 441 | 262 | 183 | 133 | 85 | 52 | 37 | 33 | 24 | 15 | 14 | 10 | 6 | 1 | 1 | 1 |
| aa | aa | 75107 | 1158.2 | 49 | 0.0e+00 | 0 | 300 | 2 | 2 | 1 | 18 | 23 | 21 | 11 | 13 | 20 | 31 | 47 | 60 | 101 | 137 | 180 | 268 | 535 | 739 | 1133 | 1725 | 3045 | 5201 | 6312 | 6436 | 6209 | 6332 | 6083 | 6187 | 6170 | 5991 | 4653 | 2729 | 1616 | 987 | 687 | 457 | 266 | 207 | 137 | 95 | 75 | 55 | 22 | 24 | 22 | 14 | 16 | 3 | 8 | 3 | 0 |
| ta | ta | 66319 | 758.9 | 49 | 0.0e+00 | 0 | 300 | 3 | 1 | 5 | 14 | 16 | 18 | 16 | 18 | 29 | 27 | 46 | 69 | 93 | 135 | 173 | 247 | 441 | 751 | 1106 | 1780 | 2911 | 5059 | 5297 | 5185 | 5354 | 5339 | 5336 | 5228 | 5084 | 5071 | 4468 | 2634 | 1506 | 976 | 625 | 388 | 263 | 169 | 129 | 84 | 82 | 49 | 36 | 19 | 12 | 11 | 6 | 5 | 6 | 2 | 0 |
| at | at | 71080 | 729.0 | 49 | 0.0e+00 | 0 | 300 | 4 | 2 | 4 | 16 | 17 | 25 | 24 | 17 | 25 | 35 | 46 | 74 | 126 | 156 | 192 | 272 | 517 | 825 | 1170 | 1861 | 3131 | 5342 | 5645 | 5563 | 5667 | 5732 | 5677 | 5682 | 5479 | 5484 | 4782 | 2803 | 1632 | 1004 | 678 | 455 | 269 | 198 | 125 | 87 | 76 | 53 | 33 | 15 | 17 | 19 | 12 | 7 | 5 | 1 | 3 |
| ag | ag | 27710 | 331.5 | 49 | 1.4e-43 | 2.2e-42 | 41.66 | 5 | 3 | 6 | 6 | 8 | 4 | 13 | 10 | 10 | 15 | 16 | 38 | 31 | 46 | 75 | 129 | 168 | 276 | 426 | 694 | 1176 | 1922 | 2357 | 2265 | 2149 | 2240 | 2302 | 2162 | 2211 | 2248 | 1850 | 1037 | 656 | 423 | 244 | 148 | 102 | 75 | 44 | 42 | 25 | 8 | 15 | 8 | 8 | 5 | 2 | 5 | 4 | 3 | 0 |
| ca | ca | 29843 | 327.1 | 49 | 9.1e-43 | 1.5e-41 | 40.84 | 6 | 3 | 3 | 6 | 12 | 7 | 10 | 8 | 3 | 19 | 15 | 40 | 39 | 54 | 82 | 144 | 220 | 271 | 522 | 745 | 1269 | 2056 | 2474 | 2429 | 2369 | 2398 | 2434 | 2358 | 2376 | 2441 | 1941 | 1122 | 660 | 417 | 311 | 212 | 112 | 90 | 55 | 35 | 20 | 11 | 11 | 11 | 8 | 7 | 4 | 7 | 1 | 0 | 1 |
| ct | ct | 28429 | 319.1 | 49 | 2.8e-41 | 4.4e-40 | 39.35 | 7 | 0 | 4 | 3 | 4 | 3 | 2 | 11 | 14 | 16 | 19 | 29 | 37 | 75 | 78 | 98 | 165 | 281 | 476 | 755 | 1206 | 2012 | 2338 | 2234 | 2325 | 2224 | 2230 | 2264 | 2319 | 2340 | 1878 | 1087 | 648 | 414 | 291 | 172 | 126 | 58 | 60 | 27 | 28 | 17 | 19 | 16 | 9 | 7 | 3 | 5 | 2 | 0 | 0 |
| tg | tg | 30375 | 274.5 | 49 | 4.0e-33 | 6.4e-32 | 31.19 | 8 | 2 | 1 | 4 | 4 | 9 | 11 | 9 | 8 | 16 | 27 | 37 | 55 | 64 | 84 | 119 | 204 | 330 | 459 | 785 | 1285 | 2141 | 2469 | 2491 | 2443 | 2414 | 2402 | 2477 | 2431 | 2404 | 1963 | 1164 | 735 | 445 | 285 | 182 | 125 | 86 | 61 | 35 | 27 | 17 | 13 | 12 | 9 | 9 | 10 | 6 | 3 | 1 | 2 |
| tc | tc | 26751 | 262.3 | 49 | 6.3e-31 | 1e-29 | 28.99 | 9 | 2 | 4 | 4 | 8 | 7 | 5 | 10 | 11 | 20 | 14 | 26 | 43 | 62 | 81 | 119 | 173 | 275 | 462 | 644 | 1140 | 1821 | 2181 | 2089 | 2180 | 2128 | 2132 | 2153 | 2165 | 2228 | 1772 | 989 | 630 | 382 | 267 | 184 | 120 | 67 | 47 | 24 | 20 | 17 | 12 | 9 | 6 | 8 | 3 | 5 | 0 | 0 | 2 |
| ga | ga | 26455 | 238.2 | 49 | 1.1e-26 | 1.8e-25 | 24.74 | 10 | 2 | 5 | 5 | 4 | 13 | 23 | 7 | 7 | 17 | 15 | 34 | 51 | 55 | 76 | 128 | 183 | 275 | 417 | 671 | 1153 | 1846 | 2234 | 2093 | 2052 | 2104 | 2096 | 2153 | 2098 | 2162 | 1716 | 985 | 630 | 389 | 239 | 156 | 100 | 79 | 44 | 43 | 24 | 10 | 9 | 11 | 12 | 9 | 7 | 3 | 4 | 4 | 2 |
| gt | gt | 24280 | 237.0 | 49 | 1.8e-26 | 2.9e-25 | 24.54 | 11 | 4 | 1 | 3 | 5 | 6 | 6 | 10 | 8 | 15 | 20 | 35 | 31 | 42 | 59 | 123 | 158 | 275 | 392 | 638 | 1044 | 1763 | 1986 | 1983 | 1951 | 1945 | 1990 | 1879 | 1874 | 1886 | 1562 | 943 | 580 | 381 | 232 | 127 | 102 | 69 | 43 | 36 | 23 | 14 | 11 | 7 | 2 | 4 | 4 | 4 | 1 | 2 | 1 |
| ac | ac | 24126 | 199.7 | 49 | 4.3e-20 | 6.8e-19 | 18.17 | 12 | 1 | 1 | 5 | 7 | 7 | 8 | 7 | 3 | 13 | 13 | 35 | 26 | 51 | 63 | 114 | 161 | 237 | 436 | 654 | 1033 | 1710 | 1980 | 1934 | 1903 | 1883 | 1915 | 1852 | 1890 | 1902 | 1619 | 961 | 561 | 352 | 269 | 168 | 103 | 72 | 57 | 36 | 26 | 10 | 11 | 15 | 7 | 4 | 3 | 3 | 2 | 3 | 0 |
| gc | gc | 9460 | 105.6 | 49 | 5.0e-06 | 7.9e-05 | 4.10 | 13 | 4 | 4 | 3 | 6 | 5 | 6 | 4 | 7 | 6 | 8 | 9 | 14 | 23 | 30 | 29 | 62 | 89 | 133 | 243 | 356 | 653 | 801 | 801 | 754 | 767 | 720 | 751 | 791 | 785 | 596 | 375 | 210 | 127 | 82 | 65 | 35 | 26 | 24 | 7 | 9 | 7 | 9 | 4 | 2 | 2 | 6 | 5 | 3 | 2 | 0 |
| cc | cc | 10820 | 96.1 | 49 | 6.8e-05 | 0.0011 | 2.96 | 14 | 4 | 5 | 5 | 2 | 9 | 7 | 8 | 8 | 11 | 5 | 8 | 14 | 25 | 32 | 52 | 73 | 106 | 191 | 272 | 472 | 769 | 879 | 888 | 847 | 879 | 818 | 854 | 882 | 897 | 674 | 377 | 263 | 138 | 118 | 51 | 58 | 32 | 23 | 14 | 11 | 9 | 8 | 3 | 5 | 3 | 1 | 1 | 4 | 2 | 3 |
| gg | gg | 10804 | 94.6 | 49 | 9.9e-05 | 0.0016 | 2.80 | 15 | 9 | 3 | 1 | 5 | 9 | 14 | 6 | 6 | 11 | 6 | 11 | 26 | 31 | 42 | 61 | 56 | 126 | 165 | 249 | 417 | 710 | 818 | 902 | 873 | 871 | 884 | 800 | 877 | 901 | 695 | 442 | 236 | 173 | 109 | 91 | 51 | 35 | 28 | 15 | 7 | 3 | 5 | 6 | 1 | 5 | 4 | 1 | 1 | 3 | 3 |
| cg | cg | 1976 | 42.8 | 49 | 7.2e-01 | 12 | -1.06 | 16 | 4 | 3 | 1 | 4 | 8 | 10 | 2 | 6 | 5 | 3 | 4 | 5 | 7 | 12 | 15 | 18 | 37 | 39 | 61 | 73 | 123 | 171 | 151 | 145 | 154 | 120 | 131 | 129 | 159 | 117 | 76 | 50 | 38 | 15 | 20 | 12 | 12 | 4 | 8 | 3 | 6 | 1 | 1 | 1 | 0 | 5 | 1 | 2 | 3 | 1 |
Host name rsat Job started 2025-12-18.171607 Job done 2025-12-18.171610 Seconds 3.74 user 3.74 system 0.01 cuser 0 ; csystem 0