position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/01/05/peak-motifs.2026-01-05.145905_2026-01-05.145905_RyN7XF/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title oxLDL_CD4+_T_cell_chromatin_accessibility -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/01/05/peak-motifs.2026-01-05.145905_2026-01-05.145905_RyN7XF/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/01/05/peak-motifs.2026-01-05.145905_2026-01-05.145905_RyN7XF/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/01/05/peak-motifs.2026-01-05.145905_2026-01-05.145905_RyN7XF/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 15572 Sum of sequence lengths 4867484 Min sequence length 0 Max sequence length 1000 Average sequence length 312 Possible positions 4851912 Oligonucleotide statistics: Total occurrences 4641711 Position interval parameters: Position interval 20 Number of windows 50 Total positions 4851912 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 40 671 2 -24 [-479,-460] -469.5 59 999 3 -23 [-459,-440] -449.5 85 1400 4 -22 [-439,-420] -429.5 108 1930 5 -21 [-419,-400] -409.5 148 2556 6 -20 [-399,-380] -389.5 203 3528 7 -19 [-379,-360] -369.5 269 4798 8 -18 [-359,-340] -349.5 403 6819 9 -17 [-339,-320] -329.5 568 9660 10 -16 [-319,-300] -309.5 827 13868 11 -15 [-299,-280] -289.5 1255 21124 12 -14 [-279,-260] -269.5 1748 30312 13 -13 [-259,-240] -249.5 2376 41425 14 -12 [-239,-220] -229.5 3008 54169 15 -11 [-219,-200] -209.5 3819 68648 16 -10 [-199,-180] -189.5 4664 85243 17 -9 [-179,-160] -169.5 5762 104550 18 -8 [-159,-140] -149.5 7867 134901 19 -7 [-139,-120] -129.5 9653 177318 20 -6 [-119,-100] -109.5 11303 210948 21 -5 [-99,-80] -89.5 13144 244986 22 -4 [-79,-60] -69.5 15511 287219 23 -3 [-59,-40] -49.5 15572 311440 24 -2 [-39,-20] -29.5 15572 311440 25 -1 [-19,0] -9.5 15572 311440 26 0 [1,20] 10.5 15572 311440 27 1 [21,40] 30.5 15572 311440 28 2 [41,60] 50.5 15572 311249 29 3 [61,80] 70.5 15314 282590 30 4 [81,100] 90.5 12977 241312 31 5 [101,120] 110.5 11181 207623 32 6 [121,140] 130.5 9524 173743 33 7 [141,160] 150.5 7707 130825 34 8 [161,180] 170.5 5670 102339 35 9 [181,200] 190.5 4575 83584 36 10 [201,220] 210.5 3755 67025 37 11 [221,240] 230.5 2959 52930 38 12 [241,260] 250.5 2321 40187 39 13 [261,280] 270.5 1705 29341 40 14 [281,300] 290.5 1227 20276 41 15 [301,320] 310.5 801 13340 42 16 [321,340] 330.5 550 9335 43 17 [341,360] 350.5 396 6554 44 18 [361,380] 370.5 265 4668 45 19 [381,400] 390.5 197 3422 46 20 [401,420] 410.5 144 2476 47 21 [421,440] 430.5 105 1886 48 22 [441,460] 450.5 84 1350 49 23 [461,480] 470.5 57 966 50 24 [481,500] 490.5 39 619 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| cg | cg | 59999 | 758.8 | 49 | 0.0e+00 | 0 | 300 | 1 | 5 | 11 | 11 | 10 | 25 | 33 | 48 | 81 | 93 | 140 | 208 | 262 | 356 | 497 | 660 | 848 | 1072 | 1553 | 2041 | 2640 | 3009 | 3776 | 4187 | 4148 | 4150 | 4254 | 4145 | 4323 | 3793 | 3034 | 2620 | 2038 | 1453 | 1115 | 801 | 677 | 559 | 384 | 305 | 191 | 137 | 97 | 68 | 45 | 38 | 26 | 13 | 8 | 3 | 8 |
| ta | ta | 264867 | 541.7 | 49 | 0.0e+00 | 0 | 300 | 2 | 49 | 52 | 93 | 112 | 157 | 209 | 264 | 408 | 556 | 786 | 1247 | 1810 | 2495 | 3150 | 4012 | 5029 | 6123 | 7793 | 9785 | 11716 | 13211 | 15290 | 16462 | 16484 | 16498 | 16279 | 16363 | 16575 | 14728 | 12987 | 11219 | 9549 | 7319 | 5924 | 4920 | 4049 | 3241 | 2315 | 1742 | 1256 | 810 | 522 | 365 | 274 | 194 | 152 | 109 | 94 | 54 | 36 |
| tt | tt | 407806 | 538.2 | 49 | 0.0e+00 | 0 | 300 | 3 | 55 | 77 | 92 | 150 | 206 | 268 | 402 | 566 | 948 | 1250 | 1849 | 2645 | 3615 | 4926 | 6223 | 7473 | 9011 | 11360 | 14830 | 17167 | 19910 | 23515 | 25395 | 25701 | 25829 | 25723 | 26089 | 25952 | 23316 | 19936 | 17335 | 14796 | 11182 | 9238 | 7674 | 6155 | 4926 | 3688 | 2523 | 1812 | 1130 | 824 | 601 | 399 | 334 | 224 | 178 | 131 | 110 | 67 |
| aa | aa | 406999 | 446.1 | 49 | 0.0e+00 | 0 | 300 | 4 | 63 | 74 | 123 | 204 | 192 | 319 | 406 | 583 | 826 | 1231 | 1919 | 2731 | 3723 | 4825 | 6095 | 7646 | 9456 | 11801 | 15207 | 17921 | 20460 | 23552 | 25844 | 25922 | 25926 | 25578 | 25629 | 25487 | 22777 | 19669 | 17025 | 14251 | 10953 | 8828 | 7356 | 5796 | 4597 | 3659 | 2625 | 1783 | 1197 | 772 | 590 | 385 | 283 | 252 | 167 | 137 | 97 | 57 |
| at | at | 308370 | 368.5 | 49 | 1.5e-50 | 2.4e-49 | 48.62 | 5 | 49 | 71 | 90 | 144 | 182 | 221 | 305 | 481 | 634 | 954 | 1447 | 2058 | 2882 | 3638 | 4708 | 5781 | 6936 | 8901 | 11399 | 13319 | 15454 | 18014 | 19367 | 19401 | 19413 | 19387 | 19312 | 19243 | 17392 | 15011 | 13000 | 11044 | 8434 | 6806 | 5610 | 4565 | 3626 | 2673 | 1930 | 1474 | 906 | 622 | 440 | 317 | 212 | 182 | 125 | 107 | 60 | 43 |
| gc | gc | 217091 | 302.3 | 49 | 3.5e-38 | 5.5e-37 | 36.26 | 6 | 27 | 37 | 54 | 68 | 108 | 149 | 185 | 280 | 425 | 565 | 868 | 1224 | 1697 | 2297 | 2759 | 3611 | 4497 | 5856 | 7873 | 9566 | 11149 | 13202 | 14182 | 14274 | 14186 | 14226 | 14227 | 14303 | 13090 | 10853 | 9434 | 7762 | 5738 | 4347 | 3418 | 2740 | 2264 | 1625 | 1219 | 867 | 518 | 429 | 277 | 200 | 141 | 94 | 73 | 47 | 41 | 19 |
| gg | gg | 248900 | 204.2 | 49 | 7.7e-21 | 1.2e-19 | 18.91 | 7 | 28 | 36 | 54 | 94 | 120 | 207 | 244 | 313 | 393 | 636 | 1029 | 1456 | 2078 | 2485 | 3310 | 4217 | 5117 | 6812 | 9061 | 10968 | 12545 | 15112 | 16141 | 16120 | 16097 | 16226 | 16345 | 16296 | 14840 | 12472 | 10535 | 8946 | 6475 | 5088 | 4157 | 3383 | 2536 | 2086 | 1483 | 1032 | 707 | 513 | 323 | 232 | 172 | 131 | 91 | 76 | 57 | 25 |
| cc | cc | 248662 | 182.3 | 49 | 3.1e-17 | 5e-16 | 15.30 | 8 | 27 | 59 | 57 | 78 | 135 | 180 | 232 | 343 | 474 | 651 | 1069 | 1598 | 2105 | 2781 | 3502 | 4219 | 5239 | 6778 | 9158 | 10751 | 12687 | 15262 | 16707 | 15918 | 15914 | 16062 | 15935 | 15979 | 14867 | 12389 | 10632 | 8687 | 6569 | 5046 | 4001 | 3321 | 2660 | 1953 | 1463 | 927 | 693 | 411 | 316 | 253 | 172 | 139 | 123 | 57 | 41 | 42 |
| tg | tg | 353303 | 157.9 | 49 | 2.2e-13 | 3.6e-12 | 11.45 | 9 | 38 | 62 | 79 | 123 | 169 | 241 | 337 | 458 | 649 | 999 | 1469 | 2192 | 2896 | 3769 | 4855 | 6067 | 7299 | 9584 | 12782 | 15232 | 17731 | 21015 | 22644 | 22840 | 23021 | 23067 | 23006 | 22817 | 21151 | 17853 | 15356 | 12767 | 9605 | 7343 | 5835 | 4755 | 3634 | 2784 | 2082 | 1498 | 925 | 698 | 453 | 344 | 270 | 157 | 146 | 97 | 67 | 42 |
| gt | gt | 253589 | 143.2 | 49 | 3.7e-11 | 5.9e-10 | 9.23 | 10 | 27 | 42 | 74 | 83 | 116 | 188 | 248 | 345 | 452 | 712 | 1053 | 1567 | 2064 | 2645 | 3455 | 4267 | 5243 | 6886 | 9054 | 11029 | 12618 | 15020 | 16329 | 16434 | 16520 | 16670 | 16437 | 16442 | 15178 | 12916 | 11155 | 9150 | 6777 | 5213 | 4249 | 3447 | 2651 | 1985 | 1530 | 992 | 689 | 475 | 346 | 237 | 202 | 123 | 99 | 76 | 52 | 27 |
| ca | ca | 353978 | 120.7 | 49 | 5.6e-08 | 9e-07 | 6.05 | 11 | 45 | 73 | 93 | 135 | 189 | 255 | 339 | 504 | 678 | 1023 | 1526 | 2167 | 2947 | 3879 | 4792 | 5942 | 7487 | 9643 | 12874 | 15242 | 18075 | 21144 | 23016 | 22928 | 22972 | 22977 | 22880 | 22935 | 20924 | 17765 | 15406 | 12801 | 9397 | 7148 | 5845 | 4691 | 3728 | 2805 | 2097 | 1430 | 922 | 675 | 488 | 329 | 235 | 192 | 133 | 91 | 68 | 48 |
| ac | ac | 253614 | 117.7 | 49 | 1.4e-07 | 2.2e-06 | 5.65 | 12 | 29 | 49 | 59 | 87 | 139 | 181 | 265 | 350 | 470 | 691 | 1121 | 1553 | 2075 | 2722 | 3399 | 4217 | 5365 | 7018 | 9257 | 10830 | 12904 | 15047 | 16445 | 16326 | 16402 | 16364 | 16398 | 16559 | 15033 | 12981 | 11031 | 9125 | 6900 | 5172 | 4200 | 3302 | 2683 | 2019 | 1525 | 1039 | 703 | 458 | 339 | 216 | 180 | 139 | 96 | 61 | 48 | 42 |
| tc | tc | 287787 | 98.6 | 49 | 3.5e-05 | 0.00055 | 3.26 | 13 | 42 | 57 | 80 | 100 | 149 | 213 | 293 | 417 | 590 | 847 | 1245 | 1821 | 2381 | 3158 | 4012 | 4895 | 6025 | 7750 | 10295 | 12177 | 14476 | 17124 | 18703 | 18815 | 18598 | 18745 | 18773 | 18529 | 16990 | 14445 | 12427 | 10325 | 7822 | 5983 | 4902 | 3875 | 3086 | 2264 | 1604 | 1135 | 786 | 538 | 366 | 279 | 199 | 151 | 115 | 83 | 53 | 49 |
| ag | ag | 344736 | 87.7 | 49 | 5.7e-04 | 0.0091 | 2.04 | 14 | 52 | 59 | 120 | 136 | 180 | 267 | 318 | 430 | 658 | 957 | 1487 | 2069 | 2871 | 3758 | 4684 | 5898 | 7339 | 9567 | 12522 | 15090 | 17511 | 20548 | 22324 | 22399 | 22223 | 22302 | 22124 | 22447 | 20174 | 17236 | 14729 | 12426 | 9217 | 7026 | 5877 | 4553 | 3687 | 2887 | 2094 | 1355 | 918 | 688 | 469 | 339 | 222 | 170 | 130 | 96 | 72 | 31 |
| ga | ga | 287783 | 83.5 | 49 | 1.6e-03 | 0.025 | 1.61 | 15 | 40 | 56 | 78 | 126 | 152 | 197 | 287 | 345 | 530 | 821 | 1238 | 1728 | 2377 | 3138 | 4006 | 4995 | 6025 | 8002 | 10533 | 12450 | 14531 | 17184 | 18769 | 18584 | 18701 | 18710 | 18544 | 18827 | 16824 | 14430 | 12167 | 10211 | 7723 | 5897 | 4862 | 3780 | 2988 | 2407 | 1734 | 1170 | 798 | 573 | 382 | 267 | 184 | 136 | 113 | 75 | 59 | 29 |
| ct | ct | 344227 | 82.2 | 49 | 2.1e-03 | 0.033 | 1.48 | 16 | 51 | 57 | 91 | 113 | 177 | 255 | 347 | 460 | 735 | 955 | 1475 | 2165 | 2915 | 3762 | 4752 | 5901 | 7332 | 9462 | 12470 | 14787 | 17346 | 20484 | 22246 | 22306 | 22091 | 22071 | 22318 | 22263 | 20273 | 17358 | 14834 | 12416 | 9451 | 7288 | 5758 | 4602 | 3673 | 2678 | 2001 | 1415 | 947 | 643 | 414 | 325 | 247 | 160 | 148 | 94 | 64 | 51 |
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