position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/01/05/peak-motifs.2026-01-05.150634_2026-01-05.150634_YIpuAE/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title 'Top_500___distal_intergenic_(ZBTB24_vs._Empty)' -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/01/05/peak-motifs.2026-01-05.150634_2026-01-05.150634_YIpuAE/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/01/05/peak-motifs.2026-01-05.150634_2026-01-05.150634_YIpuAE/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/01/05/peak-motifs.2026-01-05.150634_2026-01-05.150634_YIpuAE/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 500 Sum of sequence lengths 156785 Min sequence length 0 Max sequence length 1000 Average sequence length 313 Possible positions 156285 Oligonucleotide statistics: Total occurrences 138056 Position interval parameters: Position interval 20 Number of windows 50 Total positions 156285 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 15 276 2 -24 [-479,-460] -469.5 16 318 3 -23 [-459,-440] -449.5 20 347 4 -22 [-439,-420] -429.5 26 465 5 -21 [-419,-400] -409.5 30 568 6 -20 [-399,-380] -389.5 36 643 7 -19 [-379,-360] -369.5 39 749 8 -18 [-359,-340] -349.5 44 813 9 -17 [-339,-320] -329.5 49 918 10 -16 [-319,-300] -309.5 58 1056 11 -15 [-299,-280] -289.5 64 1221 12 -14 [-279,-260] -269.5 73 1359 13 -13 [-259,-240] -249.5 78 1485 14 -12 [-239,-220] -229.5 90 1653 15 -11 [-219,-200] -209.5 116 2128 16 -10 [-199,-180] -189.5 139 2543 17 -9 [-179,-160] -169.5 161 2995 18 -8 [-159,-140] -149.5 193 3574 19 -7 [-139,-120] -129.5 223 4117 20 -6 [-119,-100] -109.5 272 4955 21 -5 [-99,-80] -89.5 403 6773 22 -4 [-79,-60] -69.5 500 9674 23 -3 [-59,-40] -49.5 500 10000 24 -2 [-39,-20] -29.5 500 10000 25 -1 [-19,0] -9.5 500 10000 26 0 [1,20] 10.5 500 10000 27 1 [21,40] 30.5 500 10000 28 2 [41,60] 50.5 500 10000 29 3 [61,80] 70.5 500 9475 30 4 [81,100] 90.5 391 6514 31 5 [101,120] 110.5 266 4854 32 6 [121,140] 130.5 217 4064 33 7 [141,160] 150.5 189 3502 34 8 [161,180] 170.5 159 2950 35 9 [181,200] 190.5 137 2502 36 10 [201,220] 210.5 116 2075 37 11 [221,240] 230.5 87 1632 38 12 [241,260] 250.5 77 1480 39 13 [261,280] 270.5 73 1347 40 14 [281,300] 290.5 64 1206 41 15 [301,320] 310.5 56 1037 42 16 [321,340] 330.5 49 909 43 17 [341,360] 350.5 44 806 44 18 [361,380] 370.5 39 739 45 19 [381,400] 390.5 34 630 46 20 [401,420] 410.5 29 561 47 21 [421,440] 430.5 26 455 48 22 [441,460] 450.5 20 342 49 23 [461,480] 470.5 16 315 50 24 [481,500] 490.5 15 260 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| gg | gg | 9272 | 213.1 | 49 | 2.4e-22 | 3.9e-21 | 20.41 | 1 | 15 | 12 | 17 | 23 | 21 | 27 | 40 | 48 | 29 | 33 | 56 | 63 | 68 | 66 | 98 | 151 | 192 | 204 | 242 | 345 | 491 | 668 | 637 | 606 | 576 | 627 | 597 | 617 | 627 | 399 | 289 | 251 | 227 | 175 | 139 | 109 | 74 | 58 | 62 | 33 | 51 | 31 | 50 | 22 | 21 | 28 | 25 | 10 | 14 | 8 |
| cc | cc | 9694 | 198.2 | 49 | 7.7e-20 | 1.2e-18 | 17.91 | 2 | 16 | 18 | 20 | 18 | 24 | 38 | 47 | 45 | 43 | 42 | 53 | 47 | 69 | 77 | 102 | 127 | 164 | 179 | 262 | 292 | 460 | 656 | 710 | 685 | 630 | 631 | 617 | 612 | 678 | 459 | 323 | 280 | 240 | 182 | 171 | 124 | 66 | 75 | 47 | 63 | 46 | 39 | 49 | 45 | 33 | 28 | 15 | 10 | 20 | 17 |
| tt | tt | 11356 | 158.0 | 49 | 2.2e-13 | 3.5e-12 | 11.46 | 3 | 17 | 23 | 19 | 51 | 23 | 57 | 45 | 54 | 70 | 92 | 99 | 100 | 108 | 110 | 135 | 147 | 171 | 224 | 257 | 380 | 476 | 616 | 775 | 772 | 805 | 702 | 814 | 814 | 690 | 445 | 302 | 256 | 248 | 187 | 162 | 149 | 127 | 135 | 118 | 80 | 97 | 84 | 63 | 56 | 58 | 41 | 27 | 27 | 28 | 20 |
| aa | aa | 11212 | 120.9 | 49 | 5.2e-08 | 8.4e-07 | 6.08 | 4 | 11 | 20 | 32 | 40 | 49 | 43 | 57 | 54 | 71 | 102 | 96 | 124 | 108 | 116 | 128 | 196 | 218 | 287 | 299 | 340 | 472 | 684 | 802 | 782 | 698 | 756 | 738 | 716 | 611 | 393 | 315 | 262 | 226 | 181 | 175 | 131 | 130 | 91 | 108 | 93 | 74 | 69 | 55 | 74 | 37 | 29 | 48 | 21 | 23 | 27 |
| gc | gc | 7339 | 118.4 | 49 | 1.1e-07 | 1.8e-06 | 5.74 | 5 | 13 | 11 | 9 | 18 | 23 | 24 | 29 | 30 | 30 | 42 | 41 | 56 | 51 | 54 | 80 | 110 | 114 | 149 | 176 | 247 | 363 | 513 | 492 | 482 | 483 | 489 | 514 | 499 | 469 | 330 | 246 | 196 | 183 | 144 | 132 | 77 | 57 | 46 | 46 | 43 | 30 | 35 | 37 | 27 | 23 | 25 | 16 | 14 | 11 | 10 |
| ct | ct | 10924 | 102.1 | 49 | 1.3e-05 | 0.00021 | 3.68 | 6 | 19 | 23 | 17 | 29 | 46 | 41 | 53 | 56 | 56 | 57 | 73 | 75 | 79 | 106 | 137 | 142 | 180 | 227 | 284 | 349 | 495 | 720 | 737 | 690 | 796 | 715 | 737 | 681 | 717 | 497 | 365 | 288 | 237 | 206 | 178 | 125 | 85 | 94 | 72 | 83 | 65 | 60 | 49 | 39 | 39 | 32 | 18 | 22 | 13 | 20 |
| tg | tg | 10185 | 88.0 | 49 | 5.3e-04 | 0.0085 | 2.07 | 7 | 18 | 17 | 18 | 31 | 30 | 45 | 42 | 44 | 55 | 57 | 72 | 74 | 92 | 77 | 118 | 158 | 171 | 230 | 243 | 344 | 491 | 702 | 666 | 669 | 718 | 713 | 670 | 659 | 609 | 421 | 288 | 274 | 226 | 181 | 155 | 119 | 92 | 89 | 68 | 55 | 48 | 61 | 57 | 46 | 40 | 45 | 25 | 26 | 18 | 18 |
| at | at | 7854 | 86.6 | 49 | 7.5e-04 | 0.012 | 1.92 | 8 | 10 | 11 | 21 | 25 | 30 | 34 | 35 | 49 | 58 | 71 | 48 | 73 | 65 | 84 | 87 | 112 | 157 | 182 | 182 | 223 | 331 | 483 | 494 | 576 | 543 | 516 | 536 | 527 | 432 | 269 | 238 | 187 | 167 | 138 | 106 | 99 | 97 | 76 | 66 | 69 | 60 | 62 | 48 | 49 | 30 | 27 | 21 | 20 | 16 | 14 |
| cg | cg | 1931 | 82.4 | 49 | 2.0e-03 | 0.031 | 1.50 | 9 | 1 | 2 | 0 | 5 | 2 | 6 | 8 | 13 | 11 | 6 | 15 | 11 | 8 | 22 | 20 | 29 | 28 | 61 | 54 | 74 | 90 | 146 | 120 | 134 | 116 | 137 | 134 | 126 | 133 | 89 | 71 | 46 | 46 | 38 | 30 | 20 | 17 | 5 | 12 | 3 | 6 | 4 | 7 | 5 | 4 | 6 | 3 | 1 | 6 | 0 |
| tc | tc | 8993 | 79.2 | 49 | 4.0e-03 | 0.065 | 1.19 | 10 | 15 | 18 | 18 | 20 | 38 | 37 | 44 | 49 | 51 | 45 | 65 | 61 | 63 | 95 | 110 | 121 | 145 | 191 | 258 | 276 | 434 | 606 | 597 | 569 | 608 | 602 | 568 | 590 | 594 | 392 | 284 | 241 | 191 | 166 | 111 | 102 | 79 | 73 | 73 | 72 | 62 | 52 | 53 | 34 | 35 | 32 | 11 | 18 | 12 | 12 |
| ta | ta | 6392 | 66.7 | 49 | 4.7e-02 | 0.75 | 0.12 | 11 | 8 | 14 | 11 | 23 | 25 | 25 | 34 | 50 | 46 | 52 | 47 | 61 | 54 | 75 | 76 | 93 | 133 | 156 | 158 | 184 | 238 | 378 | 404 | 461 | 450 | 423 | 419 | 427 | 358 | 236 | 192 | 143 | 125 | 122 | 93 | 96 | 66 | 60 | 59 | 57 | 54 | 44 | 29 | 32 | 30 | 17 | 20 | 13 | 9 | 12 |
| ag | ag | 10528 | 65.6 | 49 | 5.7e-02 | 0.91 | 0.04 | 12 | 15 | 15 | 18 | 30 | 46 | 41 | 48 | 53 | 49 | 69 | 86 | 86 | 95 | 92 | 126 | 178 | 213 | 241 | 289 | 321 | 462 | 707 | 707 | 674 | 649 | 676 | 692 | 750 | 660 | 467 | 334 | 279 | 226 | 186 | 156 | 118 | 82 | 84 | 78 | 73 | 65 | 47 | 42 | 38 | 48 | 37 | 29 | 19 | 18 | 14 |
| gt | gt | 6801 | 65.4 | 49 | 5.8e-02 | 0.93 | 0.03 | 13 | 11 | 12 | 11 | 17 | 22 | 31 | 33 | 31 | 39 | 24 | 58 | 50 | 64 | 54 | 79 | 100 | 123 | 171 | 185 | 222 | 299 | 475 | 445 | 428 | 449 | 484 | 418 | 441 | 430 | 313 | 179 | 177 | 152 | 120 | 92 | 85 | 60 | 61 | 52 | 40 | 35 | 40 | 34 | 33 | 34 | 34 | 18 | 14 | 12 | 10 |
| ca | ca | 10282 | 64.7 | 49 | 6.6e-02 | 1.1 | -0.02 | 14 | 20 | 17 | 25 | 26 | 43 | 43 | 51 | 48 | 63 | 69 | 56 | 77 | 86 | 75 | 119 | 148 | 177 | 223 | 265 | 299 | 459 | 667 | 706 | 695 | 655 | 666 | 662 | 666 | 671 | 464 | 323 | 291 | 231 | 191 | 145 | 133 | 87 | 80 | 88 | 90 | 60 | 60 | 53 | 51 | 37 | 34 | 27 | 25 | 19 | 16 |
| ac | ac | 6799 | 63.8 | 49 | 7.6e-02 | 1.2 | -0.08 | 15 | 13 | 14 | 16 | 22 | 32 | 30 | 42 | 37 | 50 | 43 | 41 | 47 | 59 | 58 | 85 | 93 | 135 | 164 | 184 | 206 | 278 | 423 | 468 | 485 | 459 | 442 | 420 | 407 | 455 | 294 | 211 | 179 | 134 | 127 | 100 | 97 | 57 | 53 | 55 | 60 | 37 | 33 | 24 | 35 | 18 | 13 | 20 | 16 | 15 | 13 |
| ga | ga | 8494 | 58.0 | 49 | 1.8e-01 | 2.9 | -0.45 | 16 | 11 | 11 | 15 | 26 | 35 | 38 | 36 | 48 | 46 | 60 | 76 | 64 | 81 | 80 | 98 | 145 | 188 | 201 | 210 | 263 | 341 | 580 | 550 | 560 | 550 | 556 | 559 | 593 | 525 | 354 | 279 | 220 | 179 | 147 | 119 | 100 | 80 | 70 | 56 | 54 | 48 | 38 | 36 | 33 | 31 | 26 | 27 | 16 | 24 | 11 |
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