position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/01/10/peak-motifs.2026-01-10.155300_2026-01-10.155300_GuwVfF/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title promotor -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/01/10/peak-motifs.2026-01-10.155300_2026-01-10.155300_GuwVfF/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/01/10/peak-motifs.2026-01-10.155300_2026-01-10.155300_GuwVfF/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/01/10/peak-motifs.2026-01-10.155300_2026-01-10.155300_GuwVfF/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 500 Sum of sequence lengths 149801 Min sequence length 0 Max sequence length 1000 Average sequence length 299 Possible positions 149301 Oligonucleotide statistics: Total occurrences 147627 Position interval parameters: Position interval 20 Number of windows 50 Total positions 149301 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 7 110 2 -24 [-479,-460] -469.5 7 140 3 -23 [-459,-440] -449.5 8 150 4 -22 [-439,-420] -429.5 10 175 5 -21 [-419,-400] -409.5 11 220 6 -20 [-399,-380] -389.5 12 231 7 -19 [-379,-360] -369.5 17 307 8 -18 [-359,-340] -349.5 21 379 9 -17 [-339,-320] -329.5 28 478 10 -16 [-319,-300] -309.5 38 680 11 -15 [-299,-280] -289.5 46 813 12 -14 [-279,-260] -269.5 60 1103 13 -13 [-259,-240] -249.5 73 1335 14 -12 [-239,-220] -229.5 89 1632 15 -11 [-219,-200] -209.5 110 2045 16 -10 [-199,-180] -189.5 139 2475 17 -9 [-179,-160] -169.5 161 2994 18 -8 [-159,-140] -149.5 200 3639 19 -7 [-139,-120] -129.5 239 4329 20 -6 [-119,-100] -109.5 294 5276 21 -5 [-99,-80] -89.5 411 6938 22 -4 [-79,-60] -69.5 500 9699 23 -3 [-59,-40] -49.5 500 10000 24 -2 [-39,-20] -29.5 500 10000 25 -1 [-19,0] -9.5 500 10000 26 0 [1,20] 10.5 500 10000 27 1 [21,40] 30.5 500 10000 28 2 [41,60] 50.5 500 10000 29 3 [61,80] 70.5 500 9498 30 4 [81,100] 90.5 397 6710 31 5 [101,120] 110.5 288 5173 32 6 [121,140] 130.5 234 4251 33 7 [141,160] 150.5 199 3562 34 8 [161,180] 170.5 161 2948 35 9 [181,200] 190.5 135 2423 36 10 [201,220] 210.5 108 2010 37 11 [221,240] 230.5 89 1599 38 12 [241,260] 250.5 71 1310 39 13 [261,280] 270.5 59 1077 40 14 [281,300] 290.5 45 798 41 15 [301,320] 310.5 37 660 42 16 [321,340] 330.5 27 462 43 17 [341,360] 350.5 21 374 44 18 [361,380] 370.5 17 296 45 19 [381,400] 390.5 12 229 46 20 [401,420] 410.5 11 219 47 21 [421,440] 430.5 10 170 48 22 [441,460] 450.5 8 147 49 23 [461,480] 470.5 7 140 50 24 [481,500] 490.5 7 97 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| tt | tt | 8030 | 137.3 | 49 | 2.7e-10 | 4.2e-09 | 8.37 | 1 | 8 | 5 | 8 | 3 | 13 | 13 | 9 | 13 | 21 | 32 | 34 | 66 | 84 | 115 | 133 | 154 | 140 | 225 | 276 | 286 | 340 | 498 | 533 | 576 | 548 | 464 | 615 | 493 | 445 | 286 | 289 | 238 | 182 | 169 | 128 | 130 | 90 | 62 | 74 | 59 | 41 | 34 | 29 | 21 | 15 | 11 | 8 | 6 | 4 | 4 |
| gg | gg | 15031 | 136.2 | 49 | 3.9e-10 | 6.2e-09 | 8.21 | 2 | 15 | 21 | 7 | 25 | 18 | 27 | 37 | 39 | 43 | 61 | 85 | 125 | 153 | 165 | 191 | 276 | 301 | 332 | 425 | 569 | 773 | 1066 | 1048 | 1094 | 996 | 983 | 922 | 965 | 930 | 745 | 497 | 436 | 356 | 255 | 272 | 166 | 158 | 115 | 88 | 50 | 48 | 41 | 26 | 20 | 14 | 15 | 12 | 11 | 4 | 10 |
| aa | aa | 7947 | 133.2 | 49 | 1.0e-09 | 1.6e-08 | 7.79 | 3 | 1 | 5 | 12 | 16 | 22 | 17 | 20 | 29 | 24 | 49 | 50 | 64 | 78 | 105 | 132 | 152 | 199 | 238 | 247 | 264 | 334 | 483 | 523 | 529 | 540 | 522 | 558 | 524 | 440 | 316 | 251 | 186 | 165 | 150 | 130 | 110 | 83 | 86 | 71 | 46 | 34 | 29 | 16 | 19 | 14 | 14 | 18 | 7 | 14 | 11 |
| cc | cc | 14869 | 118.5 | 49 | 1.1e-07 | 1.7e-06 | 5.76 | 4 | 10 | 6 | 11 | 11 | 18 | 9 | 23 | 32 | 63 | 70 | 81 | 102 | 118 | 134 | 174 | 198 | 292 | 371 | 396 | 498 | 688 | 952 | 1026 | 942 | 1089 | 1004 | 1025 | 1017 | 1018 | 710 | 578 | 424 | 371 | 317 | 226 | 194 | 130 | 114 | 91 | 90 | 71 | 25 | 28 | 19 | 27 | 16 | 25 | 15 | 12 | 8 |
| ag | ag | 10738 | 94.9 | 49 | 9.2e-05 | 0.0015 | 2.83 | 5 | 7 | 18 | 20 | 19 | 10 | 12 | 31 | 37 | 39 | 55 | 63 | 92 | 91 | 107 | 147 | 205 | 241 | 254 | 335 | 396 | 527 | 707 | 729 | 670 | 649 | 725 | 692 | 722 | 624 | 514 | 360 | 308 | 243 | 189 | 179 | 175 | 104 | 95 | 71 | 49 | 49 | 28 | 29 | 26 | 16 | 26 | 18 | 18 | 13 | 4 |
| ta | ta | 4146 | 94.8 | 49 | 9.6e-05 | 0.0015 | 2.81 | 6 | 1 | 3 | 5 | 5 | 7 | 11 | 8 | 15 | 15 | 22 | 23 | 41 | 49 | 70 | 77 | 78 | 93 | 114 | 142 | 145 | 187 | 226 | 283 | 267 | 273 | 272 | 268 | 260 | 219 | 154 | 129 | 123 | 102 | 76 | 70 | 67 | 45 | 37 | 32 | 20 | 26 | 20 | 23 | 11 | 9 | 8 | 6 | 5 | 2 | 2 |
| at | at | 4615 | 85.8 | 49 | 8.9e-04 | 0.014 | 1.84 | 7 | 6 | 4 | 5 | 4 | 7 | 13 | 7 | 14 | 16 | 25 | 29 | 35 | 61 | 59 | 81 | 94 | 86 | 113 | 131 | 149 | 209 | 270 | 308 | 305 | 332 | 303 | 307 | 277 | 269 | 183 | 149 | 128 | 117 | 77 | 81 | 54 | 60 | 55 | 46 | 28 | 25 | 28 | 16 | 18 | 5 | 7 | 4 | 4 | 5 | 6 |
| cg | cg | 8846 | 84.2 | 49 | 1.3e-03 | 0.021 | 1.68 | 8 | 5 | 7 | 6 | 11 | 12 | 24 | 19 | 19 | 28 | 40 | 51 | 64 | 75 | 91 | 85 | 140 | 148 | 198 | 247 | 341 | 464 | 608 | 644 | 615 | 612 | 642 | 587 | 620 | 578 | 386 | 294 | 222 | 206 | 168 | 124 | 99 | 85 | 73 | 53 | 39 | 29 | 24 | 13 | 11 | 9 | 9 | 7 | 5 | 7 | 2 |
| gc | gc | 12863 | 71.1 | 49 | 2.1e-02 | 0.34 | 0.47 | 9 | 11 | 9 | 16 | 19 | 18 | 27 | 33 | 34 | 39 | 58 | 76 | 86 | 106 | 132 | 143 | 184 | 246 | 305 | 362 | 489 | 624 | 899 | 878 | 877 | 857 | 900 | 804 | 882 | 838 | 611 | 468 | 343 | 278 | 282 | 188 | 157 | 135 | 110 | 85 | 69 | 35 | 31 | 30 | 15 | 14 | 20 | 11 | 10 | 14 | 5 |
| ga | ga | 8974 | 65.5 | 49 | 5.7e-02 | 0.91 | 0.04 | 10 | 6 | 18 | 14 | 16 | 8 | 13 | 18 | 27 | 24 | 41 | 51 | 81 | 86 | 87 | 131 | 181 | 180 | 205 | 263 | 290 | 417 | 589 | 598 | 604 | 551 | 605 | 618 | 615 | 537 | 453 | 280 | 253 | 216 | 153 | 146 | 131 | 102 | 88 | 58 | 45 | 41 | 25 | 19 | 23 | 13 | 14 | 12 | 14 | 10 | 4 |
| ca | ca | 9073 | 62.6 | 49 | 9.2e-02 | 1.5 | -0.17 | 11 | 8 | 10 | 9 | 9 | 16 | 11 | 20 | 24 | 38 | 50 | 44 | 55 | 56 | 87 | 134 | 154 | 185 | 209 | 244 | 314 | 396 | 535 | 609 | 561 | 615 | 604 | 584 | 609 | 608 | 430 | 341 | 270 | 222 | 193 | 159 | 120 | 99 | 95 | 73 | 56 | 51 | 32 | 31 | 21 | 20 | 19 | 11 | 12 | 14 | 6 |
| ct | ct | 10776 | 61.0 | 49 | 1.2e-01 | 1.9 | -0.27 | 12 | 6 | 8 | 8 | 8 | 14 | 13 | 22 | 25 | 28 | 37 | 56 | 74 | 91 | 133 | 139 | 140 | 226 | 279 | 299 | 349 | 451 | 714 | 670 | 743 | 727 | 726 | 729 | 719 | 751 | 490 | 398 | 328 | 277 | 252 | 162 | 139 | 122 | 78 | 80 | 62 | 47 | 37 | 32 | 19 | 17 | 12 | 11 | 9 | 11 | 8 |
| ac | ac | 6846 | 57.3 | 49 | 1.9e-01 | 3.1 | -0.49 | 13 | 3 | 6 | 6 | 7 | 13 | 10 | 7 | 24 | 20 | 34 | 28 | 53 | 45 | 75 | 125 | 108 | 143 | 168 | 190 | 219 | 308 | 403 | 451 | 445 | 452 | 451 | 491 | 468 | 459 | 306 | 225 | 211 | 166 | 139 | 106 | 94 | 74 | 64 | 49 | 42 | 49 | 19 | 21 | 18 | 14 | 9 | 10 | 6 | 8 | 4 |
| tg | tg | 9075 | 47.3 | 49 | 5.4e-01 | 8.7 | -0.94 | 14 | 11 | 7 | 12 | 13 | 17 | 11 | 17 | 20 | 27 | 32 | 56 | 61 | 96 | 96 | 143 | 139 | 167 | 224 | 265 | 348 | 429 | 650 | 581 | 599 | 578 | 600 | 567 | 611 | 614 | 390 | 305 | 269 | 213 | 182 | 140 | 119 | 112 | 76 | 73 | 46 | 35 | 29 | 21 | 20 | 12 | 13 | 4 | 9 | 7 | 9 |
| tc | tc | 8981 | 45.8 | 49 | 6.0e-01 | 9.7 | -0.98 | 15 | 7 | 7 | 3 | 3 | 12 | 12 | 20 | 14 | 30 | 44 | 46 | 52 | 84 | 102 | 105 | 140 | 179 | 205 | 251 | 305 | 403 | 602 | 571 | 617 | 640 | 611 | 633 | 595 | 596 | 382 | 319 | 264 | 238 | 185 | 141 | 99 | 92 | 72 | 67 | 49 | 45 | 35 | 23 | 18 | 17 | 12 | 7 | 9 | 11 | 7 |
| gt | gt | 6817 | 44.2 | 49 | 6.7e-01 | 11 | -1.03 | 16 | 5 | 6 | 8 | 6 | 15 | 8 | 16 | 13 | 23 | 30 | 40 | 52 | 62 | 74 | 98 | 107 | 130 | 159 | 216 | 274 | 356 | 457 | 468 | 441 | 421 | 458 | 429 | 456 | 430 | 277 | 235 | 208 | 171 | 104 | 114 | 105 | 71 | 55 | 44 | 31 | 34 | 25 | 17 | 17 | 13 | 14 | 6 | 7 | 4 | 7 |
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