position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/01/10/peak-motifs.2026-01-10.165257_2026-01-10.165257_gp1QRU/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title Promotor -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/01/10/peak-motifs.2026-01-10.165257_2026-01-10.165257_gp1QRU/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/01/10/peak-motifs.2026-01-10.165257_2026-01-10.165257_gp1QRU/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/01/10/peak-motifs.2026-01-10.165257_2026-01-10.165257_gp1QRU/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 5379 Sum of sequence lengths 1634236 Min sequence length 0 Max sequence length 1000 Average sequence length 303 Possible positions 1628857 Oligonucleotide statistics: Total occurrences 1565971 Position interval parameters: Position interval 20 Number of windows 50 Total positions 1628857 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 109 2057 2 -24 [-479,-460] -469.5 128 2301 3 -23 [-459,-440] -449.5 146 2731 4 -22 [-439,-420] -429.5 167 3151 5 -21 [-419,-400] -409.5 185 3580 6 -20 [-399,-380] -389.5 215 4020 7 -19 [-379,-360] -369.5 256 4638 8 -18 [-359,-340] -349.5 297 5510 9 -17 [-339,-320] -329.5 355 6568 10 -16 [-319,-300] -309.5 422 7830 11 -15 [-299,-280] -289.5 512 9328 12 -14 [-279,-260] -269.5 611 11353 13 -13 [-259,-240] -249.5 757 13743 14 -12 [-239,-220] -229.5 919 16884 15 -11 [-219,-200] -209.5 1112 20439 16 -10 [-199,-180] -189.5 1369 24862 17 -9 [-179,-160] -169.5 1698 30605 18 -8 [-159,-140] -149.5 2112 38048 19 -7 [-139,-120] -129.5 2580 47038 20 -6 [-119,-100] -109.5 3301 58522 21 -5 [-99,-80] -89.5 4623 78863 22 -4 [-79,-60] -69.5 5379 104960 23 -3 [-59,-40] -49.5 5379 107580 24 -2 [-39,-20] -29.5 5379 107580 25 -1 [-19,0] -9.5 5379 107580 26 0 [1,20] 10.5 5379 107580 27 1 [21,40] 30.5 5379 107580 28 2 [41,60] 50.5 5379 107580 29 3 [61,80] 70.5 5379 103365 30 4 [81,100] 90.5 4498 76217 31 5 [101,120] 110.5 3223 57095 32 6 [121,140] 130.5 2535 46124 33 7 [141,160] 150.5 2076 37239 34 8 [161,180] 170.5 1669 29927 35 9 [181,200] 190.5 1343 24362 36 10 [201,220] 210.5 1098 20063 37 11 [221,240] 230.5 910 16563 38 12 [241,260] 250.5 748 13462 39 13 [261,280] 270.5 604 11149 40 14 [281,300] 290.5 499 9147 41 15 [301,320] 310.5 415 7699 42 16 [321,340] 330.5 351 6452 43 17 [341,360] 350.5 292 5429 44 18 [361,380] 370.5 252 4556 45 19 [381,400] 390.5 212 3957 46 20 [401,420] 410.5 184 3541 47 21 [421,440] 430.5 165 3109 48 22 [441,460] 450.5 145 2694 49 23 [461,480] 470.5 123 2267 50 24 [481,500] 490.5 109 1929 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| gg | gg | 156423 | 336.6 | 49 | 1.5e-44 | 2.4e-43 | 42.62 | 1 | 173 | 179 | 197 | 259 | 305 | 343 | 352 | 467 | 595 | 755 | 780 | 1139 | 1361 | 1566 | 1874 | 2513 | 2965 | 3830 | 4600 | 6007 | 8289 | 10444 | 10586 | 10436 | 10276 | 10356 | 10060 | 10138 | 9551 | 7450 | 5455 | 4465 | 3628 | 2834 | 2160 | 1826 | 1546 | 1171 | 965 | 805 | 734 | 574 | 503 | 411 | 326 | 284 | 270 | 259 | 208 | 153 |
| aa | aa | 86318 | 331.1 | 49 | 1.6e-43 | 2.6e-42 | 41.59 | 2 | 128 | 165 | 177 | 215 | 211 | 208 | 335 | 347 | 399 | 484 | 569 | 658 | 783 | 929 | 1124 | 1383 | 1732 | 2202 | 2446 | 3098 | 3987 | 5712 | 5857 | 5727 | 5988 | 5827 | 5777 | 5636 | 5075 | 3589 | 2867 | 2267 | 1836 | 1495 | 1253 | 1040 | 830 | 678 | 588 | 483 | 415 | 332 | 257 | 243 | 222 | 210 | 162 | 139 | 103 | 130 |
| cc | cc | 155632 | 266.9 | 49 | 9.4e-32 | 1.5e-30 | 29.82 | 3 | 183 | 188 | 229 | 279 | 336 | 318 | 399 | 450 | 604 | 676 | 849 | 981 | 1276 | 1590 | 1838 | 2202 | 3013 | 3601 | 4565 | 5488 | 7476 | 9787 | 10321 | 10070 | 9992 | 9977 | 10223 | 10641 | 10610 | 7954 | 5659 | 4469 | 3609 | 2893 | 2365 | 1912 | 1519 | 1279 | 1082 | 865 | 710 | 595 | 491 | 449 | 381 | 298 | 306 | 267 | 196 | 171 |
| tt | tt | 85694 | 250.3 | 49 | 8.5e-29 | 1.4e-27 | 26.87 | 4 | 140 | 118 | 143 | 175 | 218 | 236 | 289 | 287 | 355 | 452 | 482 | 596 | 674 | 948 | 1121 | 1246 | 1574 | 1942 | 2568 | 2968 | 3706 | 5271 | 5678 | 5708 | 5780 | 5779 | 5840 | 5615 | 5264 | 3717 | 3005 | 2441 | 1925 | 1600 | 1322 | 1124 | 1031 | 768 | 626 | 530 | 409 | 422 | 358 | 259 | 225 | 218 | 183 | 127 | 118 | 113 |
| cg | cg | 89557 | 238.9 | 49 | 8.6e-27 | 1.4e-25 | 24.86 | 5 | 72 | 73 | 124 | 121 | 150 | 217 | 191 | 250 | 301 | 357 | 434 | 590 | 709 | 883 | 1012 | 1342 | 1587 | 2052 | 2642 | 3390 | 4629 | 5944 | 6015 | 6013 | 5879 | 5915 | 5892 | 6050 | 5872 | 4306 | 3173 | 2574 | 2096 | 1589 | 1303 | 1105 | 865 | 676 | 598 | 469 | 403 | 333 | 257 | 228 | 200 | 175 | 157 | 137 | 112 | 95 |
| ta | ta | 44937 | 238.5 | 49 | 1.0e-26 | 1.6e-25 | 24.80 | 6 | 70 | 73 | 78 | 129 | 118 | 141 | 162 | 215 | 218 | 242 | 269 | 317 | 400 | 521 | 613 | 759 | 830 | 1080 | 1269 | 1479 | 1962 | 2722 | 3021 | 3019 | 3122 | 3087 | 3020 | 2895 | 2607 | 1925 | 1504 | 1281 | 979 | 784 | 709 | 555 | 489 | 399 | 364 | 274 | 222 | 198 | 175 | 133 | 117 | 111 | 95 | 67 | 63 | 55 |
| at | at | 50886 | 233.8 | 49 | 6.7e-26 | 1.1e-24 | 23.97 | 7 | 81 | 102 | 99 | 122 | 134 | 149 | 184 | 218 | 248 | 283 | 344 | 352 | 492 | 581 | 683 | 804 | 929 | 1190 | 1432 | 1739 | 2226 | 3125 | 3418 | 3381 | 3520 | 3464 | 3327 | 3266 | 3044 | 2223 | 1663 | 1455 | 1171 | 899 | 779 | 571 | 561 | 474 | 388 | 304 | 240 | 248 | 172 | 163 | 141 | 138 | 114 | 90 | 81 | 74 |
| ca | ca | 98347 | 124.5 | 49 | 1.7e-08 | 2.7e-07 | 6.58 | 8 | 143 | 194 | 190 | 196 | 232 | 231 | 312 | 342 | 437 | 488 | 626 | 705 | 787 | 1002 | 1296 | 1466 | 1849 | 2222 | 2737 | 3446 | 4582 | 6121 | 6376 | 6404 | 6420 | 6443 | 6450 | 6451 | 6381 | 4866 | 3447 | 2834 | 2330 | 1902 | 1514 | 1208 | 1023 | 869 | 706 | 559 | 449 | 423 | 336 | 289 | 223 | 227 | 196 | 148 | 150 | 119 |
| ct | ct | 114940 | 117.5 | 49 | 1.5e-07 | 2.4e-06 | 5.62 | 9 | 164 | 146 | 192 | 222 | 262 | 290 | 328 | 397 | 442 | 528 | 630 | 786 | 950 | 1182 | 1407 | 1644 | 2187 | 2651 | 3261 | 3891 | 5407 | 7221 | 7311 | 7664 | 7595 | 7550 | 7597 | 7598 | 7625 | 5554 | 4316 | 3348 | 2639 | 2221 | 1782 | 1393 | 1227 | 1001 | 786 | 659 | 546 | 442 | 384 | 307 | 290 | 232 | 200 | 185 | 150 | 150 |
| gc | gc | 133099 | 116.4 | 49 | 2.1e-07 | 3.3e-06 | 5.48 | 10 | 152 | 155 | 205 | 206 | 241 | 318 | 333 | 439 | 483 | 585 | 705 | 916 | 1090 | 1348 | 1609 | 1960 | 2448 | 3016 | 3815 | 4994 | 6719 | 8747 | 8798 | 8797 | 8623 | 8681 | 8718 | 8837 | 8604 | 6403 | 4771 | 3792 | 3056 | 2509 | 1968 | 1665 | 1362 | 1072 | 914 | 788 | 566 | 524 | 444 | 354 | 305 | 290 | 234 | 207 | 185 | 148 |
| ag | ag | 115023 | 100.3 | 49 | 2.2e-05 | 0.00035 | 3.45 | 11 | 140 | 182 | 189 | 223 | 221 | 282 | 323 | 414 | 488 | 561 | 689 | 852 | 977 | 1195 | 1509 | 1893 | 2197 | 2657 | 3351 | 4301 | 5818 | 7610 | 7547 | 7510 | 7452 | 7587 | 7637 | 7492 | 7024 | 5294 | 3842 | 3185 | 2648 | 2098 | 1745 | 1510 | 1115 | 899 | 773 | 643 | 569 | 446 | 413 | 325 | 265 | 234 | 214 | 188 | 171 | 125 |
| tg | tg | 97010 | 78.4 | 49 | 4.8e-03 | 0.077 | 1.11 | 12 | 121 | 132 | 186 | 179 | 222 | 269 | 280 | 358 | 384 | 454 | 540 | 686 | 861 | 1012 | 1244 | 1461 | 1780 | 2335 | 2871 | 3600 | 4876 | 6408 | 6337 | 6376 | 6405 | 6263 | 6309 | 6366 | 6143 | 4288 | 3439 | 2695 | 2100 | 1778 | 1473 | 1238 | 990 | 833 | 653 | 554 | 470 | 382 | 324 | 275 | 254 | 247 | 164 | 147 | 124 | 124 |
| ac | ac | 73422 | 64.3 | 49 | 7.0e-02 | 1.1 | -0.05 | 13 | 99 | 124 | 137 | 167 | 189 | 180 | 226 | 250 | 306 | 367 | 429 | 517 | 591 | 735 | 960 | 1103 | 1389 | 1711 | 2105 | 2509 | 3495 | 4575 | 4774 | 4823 | 4893 | 4908 | 4845 | 4928 | 4829 | 3491 | 2518 | 2140 | 1654 | 1406 | 1097 | 890 | 765 | 611 | 495 | 407 | 355 | 284 | 221 | 202 | 168 | 135 | 145 | 112 | 82 | 80 |
| gt | gt | 72612 | 61.6 | 49 | 1.1e-01 | 1.7 | -0.23 | 14 | 75 | 89 | 135 | 138 | 165 | 192 | 215 | 249 | 297 | 328 | 407 | 499 | 611 | 755 | 924 | 1125 | 1341 | 1732 | 2154 | 2696 | 3587 | 4833 | 4770 | 4793 | 4808 | 4679 | 4738 | 4745 | 4598 | 3246 | 2565 | 2035 | 1635 | 1248 | 1130 | 954 | 728 | 630 | 483 | 383 | 378 | 288 | 254 | 195 | 191 | 177 | 124 | 103 | 94 | 93 |
| tc | tc | 96240 | 53.6 | 49 | 3.0e-01 | 4.9 | -0.69 | 15 | 141 | 139 | 160 | 183 | 214 | 235 | 293 | 307 | 384 | 457 | 600 | 639 | 825 | 992 | 1209 | 1420 | 1889 | 2236 | 2824 | 3410 | 4604 | 6138 | 6189 | 6350 | 6449 | 6355 | 6362 | 6370 | 6335 | 4486 | 3424 | 2749 | 2248 | 1720 | 1439 | 1107 | 988 | 824 | 632 | 508 | 439 | 380 | 308 | 250 | 234 | 200 | 170 | 153 | 143 | 129 |
| ga | ga | 95831 | 51.1 | 49 | 3.9e-01 | 6.2 | -0.80 | 16 | 104 | 137 | 156 | 177 | 188 | 237 | 256 | 305 | 377 | 456 | 525 | 695 | 811 | 950 | 1182 | 1550 | 1733 | 2205 | 2766 | 3438 | 4674 | 6285 | 6294 | 6363 | 6348 | 6366 | 6380 | 6311 | 6003 | 4554 | 3268 | 2725 | 2247 | 1763 | 1441 | 1273 | 938 | 748 | 617 | 530 | 496 | 364 | 309 | 274 | 235 | 188 | 185 | 165 | 130 | 109 |
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