position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/01/10/peak-motifs.2026-01-10.165328_2026-01-10.165328_VAz8tE/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title El_distaux -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/01/10/peak-motifs.2026-01-10.165328_2026-01-10.165328_VAz8tE/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/01/10/peak-motifs.2026-01-10.165328_2026-01-10.165328_VAz8tE/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/01/10/peak-motifs.2026-01-10.165328_2026-01-10.165328_VAz8tE/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 5977 Sum of sequence lengths 1821896 Min sequence length 0 Max sequence length 1000 Average sequence length 304 Possible positions 1815919 Oligonucleotide statistics: Total occurrences 1554143 Position interval parameters: Position interval 20 Number of windows 50 Total positions 1815919 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 171 3222 2 -24 [-479,-460] -469.5 179 3517 3 -23 [-459,-440] -449.5 210 3864 4 -22 [-439,-420] -429.5 244 4530 5 -21 [-419,-400] -409.5 279 5309 6 -20 [-399,-380] -389.5 323 6044 7 -19 [-379,-360] -369.5 378 7104 8 -18 [-359,-340] -349.5 430 8105 9 -17 [-339,-320] -329.5 491 9228 10 -16 [-319,-300] -309.5 561 10633 11 -15 [-299,-280] -289.5 634 11949 12 -14 [-279,-260] -269.5 743 13813 13 -13 [-259,-240] -249.5 866 16119 14 -12 [-239,-220] -229.5 1037 19054 15 -11 [-219,-200] -209.5 1210 22606 16 -10 [-199,-180] -189.5 1452 26671 17 -9 [-179,-160] -169.5 1853 32957 18 -8 [-159,-140] -149.5 2253 41326 19 -7 [-139,-120] -129.5 2706 49676 20 -6 [-119,-100] -109.5 3371 60632 21 -5 [-99,-80] -89.5 4976 82744 22 -4 [-79,-60] -69.5 5977 116141 23 -3 [-59,-40] -49.5 5977 119540 24 -2 [-39,-20] -29.5 5977 119540 25 -1 [-19,0] -9.5 5977 119540 26 0 [1,20] 10.5 5977 119540 27 1 [21,40] 30.5 5977 119540 28 2 [41,60] 50.5 5977 119540 29 3 [61,80] 70.5 5977 114007 30 4 [81,100] 90.5 4824 79579 31 5 [101,120] 110.5 3303 59353 32 6 [121,140] 130.5 2669 48781 33 7 [141,160] 150.5 2231 40511 34 8 [161,180] 170.5 1815 32172 35 9 [181,200] 190.5 1436 26188 36 10 [201,220] 210.5 1192 22262 37 11 [221,240] 230.5 1020 18733 38 12 [241,260] 250.5 854 15871 39 13 [261,280] 270.5 732 13600 40 14 [281,300] 290.5 626 11792 41 15 [301,320] 310.5 557 10484 42 16 [321,340] 330.5 485 9110 43 17 [341,360] 350.5 429 8009 44 18 [361,380] 370.5 375 6990 45 19 [381,400] 390.5 319 5955 46 20 [401,420] 410.5 277 5240 47 21 [421,440] 430.5 242 4461 48 22 [441,460] 450.5 208 3800 49 23 [461,480] 470.5 179 3498 50 24 [481,500] 490.5 168 3039 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| aa | aa | 127017 | 605.2 | 49 | 0.0e+00 | 0 | 300 | 1 | 200 | 241 | 290 | 333 | 369 | 386 | 472 | 489 | 614 | 633 | 807 | 962 | 1054 | 1302 | 1487 | 1855 | 2257 | 2933 | 3638 | 4178 | 5659 | 8770 | 9151 | 9003 | 8934 | 8600 | 8755 | 8555 | 7410 | 4954 | 3871 | 3346 | 2749 | 2090 | 1718 | 1402 | 1266 | 1021 | 912 | 749 | 609 | 497 | 511 | 466 | 341 | 315 | 300 | 190 | 203 | 170 |
| gg | gg | 103349 | 554.0 | 49 | 0.0e+00 | 0 | 300 | 2 | 189 | 168 | 228 | 214 | 245 | 307 | 370 | 438 | 456 | 560 | 614 | 776 | 871 | 952 | 1226 | 1517 | 1934 | 2293 | 2801 | 3697 | 5328 | 7430 | 7052 | 6958 | 6755 | 6696 | 6871 | 6855 | 6510 | 4664 | 3401 | 2777 | 2361 | 1653 | 1344 | 1105 | 941 | 786 | 630 | 567 | 503 | 418 | 369 | 336 | 263 | 248 | 210 | 146 | 145 | 171 |
| cc | cc | 104032 | 550.3 | 49 | 0.0e+00 | 0 | 300 | 3 | 175 | 192 | 191 | 204 | 250 | 310 | 353 | 418 | 446 | 501 | 533 | 639 | 816 | 918 | 1102 | 1425 | 1748 | 2192 | 2727 | 3437 | 5046 | 6791 | 6911 | 6933 | 6905 | 7024 | 6808 | 7078 | 7366 | 5114 | 3480 | 2861 | 2297 | 1915 | 1480 | 1243 | 933 | 770 | 736 | 612 | 582 | 491 | 449 | 352 | 306 | 246 | 209 | 202 | 161 | 154 |
| ct | ct | 121123 | 490.2 | 49 | 0.0e+00 | 0 | 300 | 4 | 209 | 231 | 216 | 265 | 312 | 342 | 387 | 490 | 571 | 650 | 711 | 758 | 951 | 1176 | 1292 | 1569 | 2063 | 2587 | 3201 | 3952 | 5539 | 7853 | 8109 | 8107 | 8214 | 8123 | 8161 | 8232 | 8488 | 5762 | 4136 | 3264 | 2691 | 2164 | 1704 | 1429 | 1157 | 974 | 781 | 788 | 617 | 575 | 427 | 429 | 332 | 289 | 230 | 228 | 192 | 195 |
| tt | tt | 127630 | 453.0 | 49 | 0.0e+00 | 0 | 300 | 5 | 215 | 200 | 238 | 308 | 312 | 408 | 398 | 519 | 557 | 728 | 809 | 922 | 1103 | 1270 | 1571 | 1748 | 2192 | 2790 | 3382 | 3999 | 5140 | 7714 | 8680 | 8951 | 8798 | 8762 | 8823 | 8968 | 8510 | 5701 | 4188 | 3403 | 2885 | 2143 | 1850 | 1471 | 1236 | 1092 | 911 | 724 | 696 | 623 | 608 | 472 | 375 | 337 | 282 | 232 | 214 | 172 |
| ag | ag | 120054 | 398.7 | 49 | 2.6e-56 | 4.1e-55 | 54.38 | 6 | 186 | 221 | 230 | 258 | 330 | 328 | 439 | 499 | 556 | 608 | 737 | 813 | 1017 | 1166 | 1430 | 1761 | 2226 | 2747 | 3310 | 4085 | 6128 | 8346 | 8086 | 8086 | 8074 | 8074 | 8026 | 8065 | 7473 | 5229 | 3849 | 3090 | 2519 | 2044 | 1611 | 1376 | 1181 | 957 | 789 | 693 | 595 | 552 | 439 | 377 | 331 | 279 | 242 | 209 | 209 | 178 |
| tg | tg | 116213 | 344.7 | 49 | 4.6e-46 | 7.3e-45 | 44.14 | 7 | 176 | 190 | 222 | 255 | 285 | 341 | 455 | 497 | 516 | 641 | 717 | 833 | 966 | 1176 | 1378 | 1635 | 2049 | 2618 | 3075 | 3991 | 5671 | 8046 | 7867 | 7787 | 7739 | 7973 | 7700 | 7875 | 7462 | 5143 | 3705 | 3065 | 2596 | 1946 | 1591 | 1281 | 1062 | 953 | 789 | 633 | 588 | 500 | 428 | 412 | 312 | 281 | 239 | 194 | 178 | 181 |
| tc | tc | 99763 | 333.0 | 49 | 7.3e-44 | 1.2e-42 | 41.93 | 8 | 179 | 206 | 178 | 234 | 266 | 284 | 298 | 400 | 455 | 550 | 587 | 661 | 791 | 971 | 1117 | 1349 | 1750 | 2191 | 2662 | 3278 | 4647 | 6491 | 6645 | 6678 | 6693 | 6734 | 6785 | 6854 | 6979 | 4510 | 3274 | 2686 | 2255 | 1706 | 1334 | 1164 | 961 | 767 | 643 | 617 | 499 | 465 | 394 | 327 | 295 | 258 | 191 | 188 | 160 | 156 |
| ca | ca | 115722 | 314.4 | 49 | 2.0e-40 | 3.3e-39 | 38.49 | 9 | 206 | 199 | 234 | 262 | 315 | 338 | 396 | 445 | 558 | 595 | 643 | 772 | 941 | 1066 | 1346 | 1593 | 1977 | 2535 | 3152 | 3802 | 5403 | 7489 | 7735 | 7751 | 7846 | 7802 | 7880 | 7831 | 7776 | 5403 | 3769 | 3132 | 2573 | 2111 | 1635 | 1302 | 1119 | 921 | 783 | 652 | 601 | 531 | 457 | 364 | 359 | 294 | 242 | 216 | 192 | 178 |
| gc | gc | 80931 | 287.5 | 49 | 1.8e-35 | 2.9e-34 | 33.54 | 10 | 130 | 155 | 151 | 186 | 203 | 227 | 308 | 325 | 403 | 433 | 441 | 543 | 621 | 739 | 946 | 1091 | 1405 | 1799 | 2171 | 2748 | 4043 | 5515 | 5419 | 5302 | 5458 | 5436 | 5325 | 5491 | 5326 | 3742 | 2702 | 2171 | 1737 | 1461 | 1150 | 898 | 757 | 657 | 536 | 455 | 405 | 375 | 297 | 292 | 222 | 171 | 162 | 151 | 126 | 124 |
| ga | ga | 98305 | 275.8 | 49 | 2.4e-33 | 3.8e-32 | 31.42 | 11 | 142 | 167 | 180 | 193 | 263 | 284 | 375 | 412 | 409 | 513 | 612 | 673 | 856 | 982 | 1176 | 1389 | 1768 | 2251 | 2584 | 3331 | 4658 | 6811 | 6654 | 6684 | 6615 | 6610 | 6604 | 6623 | 6165 | 4410 | 3285 | 2593 | 2156 | 1657 | 1328 | 1120 | 946 | 788 | 663 | 565 | 489 | 408 | 383 | 325 | 246 | 251 | 214 | 164 | 168 | 162 |
| ta | ta | 73162 | 242.6 | 49 | 1.9e-27 | 3e-26 | 25.52 | 12 | 118 | 130 | 122 | 168 | 184 | 217 | 266 | 295 | 356 | 395 | 459 | 528 | 657 | 788 | 867 | 1054 | 1249 | 1605 | 2004 | 2417 | 3178 | 4564 | 5127 | 5082 | 5254 | 5055 | 5131 | 4954 | 4472 | 2942 | 2334 | 1959 | 1536 | 1266 | 1064 | 898 | 711 | 628 | 509 | 462 | 417 | 350 | 278 | 240 | 195 | 173 | 161 | 124 | 127 | 92 |
| ac | ac | 77353 | 241.8 | 49 | 2.7e-27 | 4.3e-26 | 25.37 | 13 | 138 | 130 | 155 | 158 | 217 | 247 | 275 | 297 | 367 | 390 | 445 | 525 | 658 | 728 | 870 | 1074 | 1297 | 1681 | 2179 | 2637 | 3526 | 4940 | 5244 | 5215 | 5264 | 5219 | 5309 | 5178 | 5267 | 3530 | 2553 | 2084 | 1704 | 1400 | 1053 | 889 | 727 | 593 | 505 | 463 | 410 | 349 | 269 | 250 | 213 | 194 | 163 | 129 | 130 | 115 |
| at | at | 89958 | 239.3 | 49 | 7.2e-27 | 1.2e-25 | 24.94 | 14 | 141 | 154 | 167 | 201 | 232 | 279 | 307 | 371 | 401 | 520 | 545 | 661 | 793 | 968 | 1119 | 1234 | 1568 | 2012 | 2357 | 2956 | 3884 | 5711 | 6226 | 6288 | 6318 | 6203 | 6252 | 6167 | 5574 | 3708 | 2908 | 2400 | 1982 | 1535 | 1276 | 1057 | 863 | 755 | 625 | 544 | 497 | 378 | 384 | 304 | 248 | 235 | 207 | 168 | 139 | 136 |
| gt | gt | 78269 | 214.9 | 49 | 1.2e-22 | 1.9e-21 | 20.73 | 15 | 135 | 135 | 136 | 188 | 197 | 226 | 287 | 329 | 359 | 399 | 497 | 586 | 659 | 781 | 924 | 1157 | 1386 | 1803 | 2107 | 2638 | 3719 | 5413 | 5327 | 5276 | 5146 | 5389 | 5281 | 5248 | 5006 | 3419 | 2492 | 2057 | 1743 | 1321 | 1026 | 913 | 736 | 655 | 518 | 410 | 402 | 359 | 285 | 258 | 239 | 190 | 154 | 127 | 126 | 105 |
| cg | cg | 21262 | 114.0 | 49 | 4.3e-07 | 6.9e-06 | 5.16 | 16 | 33 | 45 | 31 | 52 | 64 | 62 | 84 | 84 | 99 | 117 | 124 | 162 | 162 | 184 | 269 | 311 | 355 | 501 | 572 | 787 | 1009 | 1437 | 1433 | 1350 | 1404 | 1408 | 1431 | 1432 | 1367 | 1007 | 781 | 577 | 480 | 373 | 267 | 230 | 187 | 158 | 136 | 105 | 102 | 81 | 85 | 75 | 65 | 47 | 46 | 34 | 35 | 22 |
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