position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/01/22/peak-motifs.2026-01-22.084309_2026-01-22.084309_ujjUO0/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title WT24h-hupA_sensitive -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/01/22/peak-motifs.2026-01-22.084309_2026-01-22.084309_ujjUO0/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/01/22/peak-motifs.2026-01-22.084309_2026-01-22.084309_ujjUO0/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/01/22/peak-motifs.2026-01-22.084309_2026-01-22.084309_ujjUO0/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 551 Sum of sequence lengths 225925 Min sequence length 0 Max sequence length 1000 Average sequence length 410 Possible positions 225374 Oligonucleotide statistics: Total occurrences 223126 Position interval parameters: Position interval 20 Number of windows 50 Total positions 225374 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 11 200 2 -24 [-479,-460] -469.5 14 259 3 -23 [-459,-440] -449.5 17 316 4 -22 [-439,-420] -429.5 20 379 5 -21 [-419,-400] -409.5 24 442 6 -20 [-399,-380] -389.5 29 518 7 -19 [-379,-360] -369.5 33 628 8 -18 [-359,-340] -349.5 42 750 9 -17 [-339,-320] -329.5 55 1004 10 -16 [-319,-300] -309.5 77 1334 11 -15 [-299,-280] -289.5 95 1738 12 -14 [-279,-260] -269.5 124 2151 13 -13 [-259,-240] -249.5 158 2832 14 -12 [-239,-220] -229.5 186 3442 15 -11 [-219,-200] -209.5 229 4199 16 -10 [-199,-180] -189.5 276 5048 17 -9 [-179,-160] -169.5 339 6198 18 -8 [-159,-140] -149.5 421 7651 19 -7 [-139,-120] -129.5 478 9007 20 -6 [-119,-100] -109.5 534 10094 21 -5 [-99,-80] -89.5 551 10957 22 -4 [-79,-60] -69.5 551 11020 23 -3 [-59,-40] -49.5 551 11020 24 -2 [-39,-20] -29.5 551 11020 25 -1 [-19,0] -9.5 551 11020 26 0 [1,20] 10.5 551 11020 27 1 [21,40] 30.5 551 11020 28 2 [41,60] 50.5 551 11020 29 3 [61,80] 70.5 551 11020 30 4 [81,100] 90.5 551 10923 31 5 [101,120] 110.5 532 9985 32 6 [121,140] 130.5 476 8898 33 7 [141,160] 150.5 414 7490 34 8 [161,180] 170.5 332 6075 35 9 [181,200] 190.5 272 4953 36 10 [201,220] 210.5 228 4114 37 11 [221,240] 230.5 182 3386 38 12 [241,260] 250.5 154 2767 39 13 [261,280] 270.5 118 2090 40 14 [281,300] 290.5 94 1702 41 15 [301,320] 310.5 76 1290 42 16 [321,340] 330.5 54 977 43 17 [341,360] 350.5 39 732 44 18 [361,380] 370.5 33 620 45 19 [381,400] 390.5 28 509 46 20 [401,420] 410.5 23 433 47 21 [421,440] 430.5 20 373 48 22 [441,460] 450.5 17 310 49 23 [461,480] 470.5 14 252 50 24 [481,500] 490.5 10 188 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| aa | aa | 3799 | 150.4 | 49 | 3.1e-12 | 4.9e-11 | 10.31 | 1 | 9 | 7 | 6 | 3 | 9 | 14 | 9 | 6 | 20 | 26 | 28 | 33 | 32 | 48 | 56 | 69 | 93 | 123 | 139 | 145 | 195 | 201 | 243 | 235 | 279 | 235 | 186 | 158 | 175 | 144 | 149 | 146 | 115 | 94 | 62 | 64 | 51 | 42 | 36 | 25 | 23 | 13 | 13 | 9 | 6 | 6 | 6 | 3 | 6 | 4 |
| tt | tt | 3994 | 108.3 | 49 | 2.3e-06 | 3.7e-05 | 4.43 | 2 | 2 | 3 | 5 | 5 | 10 | 3 | 9 | 14 | 11 | 28 | 23 | 48 | 48 | 52 | 78 | 68 | 81 | 121 | 146 | 190 | 195 | 207 | 178 | 221 | 229 | 274 | 227 | 207 | 211 | 175 | 168 | 131 | 133 | 114 | 71 | 72 | 49 | 45 | 28 | 30 | 23 | 13 | 14 | 5 | 11 | 11 | 2 | 2 | 2 | 1 |
| gg | gg | 24543 | 64.8 | 49 | 6.5e-02 | 1 | -0.02 | 3 | 25 | 29 | 37 | 42 | 48 | 54 | 68 | 76 | 109 | 153 | 177 | 230 | 359 | 387 | 457 | 584 | 684 | 815 | 977 | 1072 | 1204 | 1190 | 1143 | 1152 | 1147 | 1114 | 1247 | 1217 | 1176 | 1148 | 1095 | 972 | 845 | 722 | 571 | 430 | 436 | 308 | 213 | 173 | 139 | 111 | 90 | 77 | 70 | 41 | 43 | 36 | 33 | 17 |
| at | at | 4006 | 61.4 | 49 | 1.1e-01 | 1.8 | -0.25 | 4 | 6 | 7 | 4 | 11 | 7 | 13 | 11 | 12 | 13 | 20 | 22 | 29 | 57 | 68 | 86 | 95 | 120 | 145 | 169 | 191 | 172 | 182 | 219 | 191 | 168 | 170 | 207 | 208 | 203 | 216 | 185 | 172 | 136 | 94 | 78 | 68 | 46 | 46 | 50 | 33 | 12 | 12 | 14 | 7 | 6 | 6 | 3 | 4 | 9 | 3 |
| tc | tc | 14316 | 57.3 | 49 | 1.9e-01 | 3.1 | -0.49 | 5 | 8 | 13 | 28 | 18 | 29 | 28 | 35 | 53 | 59 | 80 | 101 | 127 | 171 | 206 | 296 | 328 | 406 | 483 | 591 | 679 | 699 | 684 | 714 | 705 | 670 | 702 | 692 | 713 | 679 | 706 | 646 | 538 | 516 | 370 | 286 | 304 | 187 | 189 | 142 | 94 | 78 | 59 | 51 | 20 | 25 | 30 | 26 | 22 | 17 | 13 |
| cc | cc | 24883 | 54.9 | 49 | 2.6e-01 | 4.2 | -0.62 | 6 | 23 | 26 | 36 | 33 | 42 | 60 | 66 | 89 | 123 | 133 | 203 | 234 | 328 | 403 | 454 | 597 | 720 | 899 | 1032 | 1163 | 1212 | 1243 | 1264 | 1171 | 1211 | 1132 | 1149 | 1192 | 1205 | 1250 | 1098 | 936 | 802 | 634 | 572 | 450 | 349 | 320 | 229 | 213 | 141 | 100 | 67 | 67 | 52 | 51 | 37 | 32 | 22 | 18 |
| ca | ca | 11305 | 53.7 | 49 | 3.0e-01 | 4.8 | -0.68 | 7 | 15 | 20 | 9 | 24 | 23 | 23 | 26 | 41 | 61 | 55 | 84 | 95 | 129 | 187 | 227 | 297 | 313 | 402 | 430 | 540 | 537 | 568 | 577 | 569 | 569 | 601 | 548 | 531 | 547 | 520 | 468 | 425 | 356 | 295 | 249 | 200 | 165 | 132 | 94 | 93 | 60 | 43 | 33 | 28 | 26 | 16 | 18 | 13 | 13 | 10 |
| tg | tg | 11740 | 52.8 | 49 | 3.3e-01 | 5.3 | -0.72 | 8 | 9 | 15 | 12 | 20 | 29 | 30 | 42 | 49 | 51 | 90 | 88 | 122 | 147 | 165 | 220 | 253 | 298 | 400 | 442 | 489 | 556 | 526 | 569 | 578 | 587 | 608 | 588 | 587 | 620 | 566 | 510 | 489 | 364 | 318 | 276 | 200 | 186 | 126 | 106 | 85 | 82 | 50 | 37 | 37 | 37 | 21 | 16 | 20 | 14 | 10 |
| ac | ac | 13420 | 45.5 | 49 | 6.2e-01 | 9.9 | -0.99 | 9 | 17 | 18 | 16 | 30 | 32 | 32 | 33 | 43 | 61 | 77 | 120 | 125 | 150 | 201 | 238 | 309 | 365 | 449 | 544 | 594 | 657 | 698 | 643 | 691 | 708 | 670 | 662 | 611 | 664 | 627 | 555 | 517 | 420 | 348 | 295 | 252 | 217 | 158 | 114 | 116 | 75 | 67 | 35 | 44 | 26 | 20 | 22 | 22 | 16 | 16 |
| ta | ta | 2215 | 40.5 | 49 | 8.0e-01 | 13 | -1.11 | 10 | 3 | 1 | 6 | 5 | 4 | 6 | 3 | 6 | 8 | 13 | 21 | 23 | 33 | 41 | 37 | 52 | 49 | 74 | 86 | 99 | 108 | 110 | 113 | 100 | 121 | 99 | 121 | 123 | 90 | 100 | 112 | 103 | 56 | 60 | 42 | 37 | 25 | 28 | 27 | 17 | 7 | 14 | 8 | 8 | 2 | 2 | 4 | 4 | 1 | 3 |
| gt | gt | 13208 | 40.2 | 49 | 8.1e-01 | 13 | -1.11 | 11 | 8 | 12 | 18 | 17 | 30 | 24 | 41 | 50 | 56 | 86 | 90 | 136 | 167 | 190 | 225 | 298 | 361 | 408 | 531 | 584 | 649 | 619 | 620 | 655 | 644 | 698 | 697 | 664 | 650 | 639 | 591 | 530 | 421 | 369 | 285 | 259 | 194 | 162 | 111 | 83 | 84 | 60 | 42 | 33 | 29 | 24 | 26 | 19 | 7 | 12 |
| gc | gc | 27196 | 39.5 | 49 | 8.3e-01 | 13 | -1.12 | 12 | 18 | 30 | 41 | 47 | 48 | 73 | 86 | 88 | 129 | 159 | 209 | 262 | 337 | 420 | 536 | 608 | 751 | 934 | 1062 | 1216 | 1335 | 1319 | 1274 | 1280 | 1259 | 1279 | 1273 | 1366 | 1316 | 1347 | 1209 | 1087 | 937 | 748 | 642 | 486 | 426 | 348 | 256 | 212 | 171 | 127 | 87 | 92 | 66 | 62 | 50 | 33 | 33 | 22 |
| ag | ag | 10476 | 37.6 | 49 | 8.8e-01 | 14 | -1.15 | 13 | 9 | 17 | 13 | 20 | 12 | 23 | 27 | 32 | 57 | 58 | 88 | 96 | 139 | 171 | 188 | 234 | 279 | 364 | 403 | 468 | 481 | 502 | 514 | 496 | 527 | 538 | 526 | 514 | 475 | 491 | 475 | 442 | 343 | 318 | 227 | 187 | 161 | 134 | 105 | 62 | 52 | 44 | 40 | 29 | 30 | 20 | 18 | 11 | 8 | 8 |
| cg | cg | 32605 | 37.3 | 49 | 8.9e-01 | 14 | -1.15 | 14 | 25 | 34 | 46 | 57 | 66 | 75 | 95 | 109 | 141 | 187 | 245 | 316 | 385 | 489 | 615 | 707 | 940 | 1076 | 1342 | 1456 | 1611 | 1605 | 1524 | 1569 | 1486 | 1527 | 1561 | 1595 | 1606 | 1608 | 1480 | 1293 | 1128 | 896 | 744 | 621 | 503 | 420 | 306 | 254 | 195 | 153 | 109 | 95 | 64 | 72 | 65 | 45 | 36 | 28 |
| ct | ct | 11047 | 33.5 | 49 | 9.6e-01 | 15 | -1.18 | 15 | 7 | 8 | 23 | 17 | 25 | 25 | 34 | 39 | 47 | 79 | 91 | 108 | 128 | 171 | 218 | 238 | 289 | 395 | 426 | 483 | 545 | 518 | 561 | 546 | 556 | 545 | 502 | 547 | 538 | 535 | 466 | 428 | 387 | 283 | 233 | 222 | 156 | 132 | 113 | 85 | 69 | 48 | 39 | 27 | 28 | 22 | 17 | 21 | 17 | 10 |
| ga | ga | 14373 | 32.9 | 49 | 9.6e-01 | 15 | -1.19 | 16 | 16 | 19 | 16 | 30 | 28 | 35 | 43 | 43 | 58 | 88 | 126 | 127 | 182 | 214 | 248 | 284 | 408 | 483 | 595 | 611 | 661 | 707 | 709 | 701 | 708 | 688 | 710 | 667 | 720 | 691 | 650 | 612 | 479 | 406 | 320 | 262 | 235 | 177 | 145 | 107 | 72 | 63 | 53 | 42 | 31 | 29 | 20 | 23 | 18 | 13 |
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