position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/01/22/peak-motifs.2026-01-22.084429_2026-01-22.084429_vEeoc0/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title WT48h_hupA_sensitive -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/01/22/peak-motifs.2026-01-22.084429_2026-01-22.084429_vEeoc0/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/01/22/peak-motifs.2026-01-22.084429_2026-01-22.084429_vEeoc0/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/01/22/peak-motifs.2026-01-22.084429_2026-01-22.084429_vEeoc0/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 493 Sum of sequence lengths 239920 Min sequence length 0 Max sequence length 1000 Average sequence length 486 Possible positions 239427 Oligonucleotide statistics: Total occurrences 236788 Position interval parameters: Position interval 20 Number of windows 50 Total positions 239427 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 28 520 2 -24 [-479,-460] -469.5 30 567 3 -23 [-459,-440] -449.5 33 635 4 -22 [-439,-420] -429.5 39 710 5 -21 [-419,-400] -409.5 44 831 6 -20 [-399,-380] -389.5 54 975 7 -19 [-379,-360] -369.5 64 1198 8 -18 [-359,-340] -349.5 73 1371 9 -17 [-339,-320] -329.5 88 1603 10 -16 [-319,-300] -309.5 111 2015 11 -15 [-299,-280] -289.5 137 2556 12 -14 [-279,-260] -269.5 171 3077 13 -13 [-259,-240] -249.5 200 3676 14 -12 [-239,-220] -229.5 240 4448 15 -11 [-219,-200] -209.5 283 5286 16 -10 [-199,-180] -189.5 342 6289 17 -9 [-179,-160] -169.5 399 7427 18 -8 [-159,-140] -149.5 447 8514 19 -7 [-139,-120] -129.5 486 9376 20 -6 [-119,-100] -109.5 493 9815 21 -5 [-99,-80] -89.5 493 9860 22 -4 [-79,-60] -69.5 493 9860 23 -3 [-59,-40] -49.5 493 9860 24 -2 [-39,-20] -29.5 493 9860 25 -1 [-19,0] -9.5 493 9860 26 0 [1,20] 10.5 493 9860 27 1 [21,40] 30.5 493 9860 28 2 [41,60] 50.5 493 9860 29 3 [61,80] 70.5 493 9860 30 4 [81,100] 90.5 493 9860 31 5 [101,120] 110.5 493 9799 32 6 [121,140] 130.5 483 9305 33 7 [141,160] 150.5 444 8403 34 8 [161,180] 170.5 392 7319 35 9 [181,200] 190.5 341 6180 36 10 [201,220] 210.5 283 5196 37 11 [221,240] 230.5 237 4373 38 12 [241,260] 250.5 198 3621 39 13 [261,280] 270.5 167 3010 40 14 [281,300] 290.5 135 2504 41 15 [301,320] 310.5 111 1971 42 16 [321,340] 330.5 87 1575 43 17 [341,360] 350.5 72 1353 44 18 [361,380] 370.5 64 1177 45 19 [381,400] 390.5 52 954 46 20 [401,420] 410.5 44 819 47 21 [421,440] 430.5 39 698 48 22 [441,460] 450.5 33 627 49 23 [461,480] 470.5 29 563 50 24 [481,500] 490.5 28 491 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| tt | tt | 4303 | 117.4 | 49 | 1.5e-07 | 2.5e-06 | 5.61 | 1 | 12 | 10 | 13 | 15 | 17 | 10 | 20 | 28 | 32 | 39 | 41 | 50 | 75 | 93 | 105 | 100 | 113 | 127 | 148 | 186 | 175 | 209 | 195 | 174 | 221 | 231 | 205 | 185 | 202 | 207 | 152 | 150 | 139 | 107 | 93 | 94 | 65 | 52 | 41 | 36 | 26 | 20 | 18 | 16 | 21 | 16 | 7 | 5 | 3 | 4 |
| aa | aa | 4137 | 102.0 | 49 | 1.4e-05 | 0.00022 | 3.66 | 2 | 15 | 10 | 12 | 15 | 7 | 21 | 22 | 29 | 20 | 28 | 47 | 61 | 59 | 72 | 89 | 79 | 113 | 116 | 174 | 169 | 173 | 173 | 205 | 206 | 206 | 211 | 207 | 165 | 165 | 131 | 166 | 158 | 122 | 106 | 105 | 74 | 81 | 57 | 50 | 35 | 35 | 27 | 33 | 21 | 18 | 13 | 7 | 10 | 9 | 10 |
| cc | cc | 26054 | 69.1 | 49 | 3.1e-02 | 0.49 | 0.31 | 3 | 55 | 60 | 66 | 69 | 73 | 115 | 152 | 149 | 170 | 228 | 281 | 347 | 430 | 473 | 539 | 696 | 888 | 947 | 1012 | 1098 | 1127 | 1082 | 1123 | 1056 | 999 | 1069 | 1043 | 1046 | 1061 | 1089 | 1097 | 999 | 868 | 774 | 706 | 616 | 457 | 394 | 316 | 266 | 190 | 142 | 136 | 121 | 90 | 87 | 64 | 83 | 45 | 60 |
| gg | gg | 25805 | 68.0 | 49 | 3.7e-02 | 0.6 | 0.23 | 4 | 54 | 64 | 66 | 89 | 121 | 97 | 136 | 148 | 176 | 242 | 295 | 315 | 373 | 495 | 609 | 673 | 823 | 893 | 1008 | 1044 | 1049 | 1058 | 1093 | 1020 | 1001 | 1053 | 1015 | 1061 | 1034 | 1057 | 1061 | 1049 | 867 | 866 | 688 | 534 | 460 | 410 | 322 | 267 | 219 | 181 | 124 | 132 | 109 | 76 | 98 | 63 | 74 | 43 |
| at | at | 4480 | 65.5 | 49 | 5.8e-02 | 0.92 | 0.04 | 5 | 11 | 12 | 21 | 16 | 20 | 18 | 22 | 20 | 44 | 43 | 43 | 59 | 66 | 77 | 105 | 139 | 137 | 166 | 169 | 189 | 179 | 186 | 178 | 197 | 176 | 172 | 145 | 191 | 181 | 182 | 178 | 207 | 162 | 132 | 116 | 100 | 52 | 62 | 71 | 46 | 28 | 29 | 25 | 24 | 14 | 19 | 15 | 11 | 17 | 8 |
| ct | ct | 11925 | 52.9 | 49 | 3.3e-01 | 5.2 | -0.72 | 6 | 26 | 22 | 28 | 38 | 32 | 49 | 51 | 65 | 97 | 98 | 124 | 164 | 213 | 231 | 246 | 324 | 369 | 439 | 496 | 476 | 504 | 494 | 483 | 495 | 502 | 483 | 447 | 490 | 499 | 486 | 447 | 430 | 404 | 354 | 311 | 271 | 205 | 179 | 158 | 144 | 109 | 89 | 71 | 58 | 52 | 38 | 38 | 28 | 26 | 42 |
| ac | ac | 14279 | 50.0 | 49 | 4.3e-01 | 6.9 | -0.84 | 7 | 32 | 30 | 38 | 41 | 42 | 53 | 77 | 82 | 84 | 106 | 143 | 190 | 226 | 245 | 305 | 372 | 430 | 515 | 567 | 582 | 595 | 594 | 594 | 588 | 613 | 593 | 643 | 555 | 599 | 573 | 601 | 512 | 511 | 419 | 334 | 307 | 284 | 206 | 180 | 142 | 115 | 94 | 106 | 92 | 55 | 55 | 38 | 43 | 45 | 33 |
| tc | tc | 15307 | 49.3 | 49 | 4.6e-01 | 7.4 | -0.87 | 8 | 32 | 34 | 41 | 32 | 49 | 64 | 77 | 85 | 123 | 114 | 135 | 200 | 265 | 292 | 325 | 429 | 489 | 561 | 602 | 660 | 626 | 620 | 622 | 643 | 646 | 602 | 612 | 604 | 637 | 614 | 603 | 564 | 541 | 489 | 412 | 348 | 267 | 230 | 213 | 181 | 102 | 106 | 75 | 65 | 64 | 61 | 45 | 37 | 34 | 35 |
| ca | ca | 12421 | 49.2 | 49 | 4.7e-01 | 7.5 | -0.87 | 9 | 33 | 35 | 33 | 34 | 48 | 58 | 76 | 70 | 76 | 96 | 132 | 172 | 198 | 228 | 282 | 350 | 399 | 444 | 467 | 470 | 516 | 521 | 509 | 501 | 514 | 513 | 547 | 492 | 486 | 525 | 511 | 440 | 444 | 359 | 307 | 272 | 239 | 182 | 159 | 119 | 118 | 73 | 86 | 71 | 51 | 48 | 19 | 33 | 38 | 27 |
| gc | gc | 28591 | 44.5 | 49 | 6.6e-01 | 10 | -1.02 | 10 | 57 | 65 | 74 | 80 | 108 | 118 | 143 | 162 | 187 | 263 | 342 | 361 | 418 | 552 | 623 | 753 | 906 | 1040 | 1100 | 1120 | 1196 | 1176 | 1121 | 1132 | 1142 | 1150 | 1111 | 1198 | 1162 | 1216 | 1189 | 1083 | 1013 | 896 | 794 | 623 | 553 | 442 | 360 | 306 | 276 | 187 | 172 | 149 | 116 | 86 | 80 | 70 | 63 | 57 |
| ta | ta | 2432 | 43.2 | 49 | 7.1e-01 | 11 | -1.05 | 11 | 6 | 7 | 5 | 7 | 12 | 10 | 10 | 11 | 18 | 24 | 19 | 34 | 37 | 40 | 62 | 56 | 64 | 104 | 91 | 102 | 105 | 98 | 97 | 105 | 97 | 102 | 93 | 123 | 106 | 90 | 93 | 97 | 95 | 70 | 47 | 53 | 44 | 39 | 21 | 22 | 20 | 17 | 12 | 12 | 8 | 6 | 13 | 8 | 11 | 9 |
| ag | ag | 11286 | 43.1 | 49 | 7.1e-01 | 11 | -1.06 | 12 | 20 | 34 | 23 | 39 | 45 | 61 | 54 | 76 | 71 | 96 | 133 | 149 | 172 | 212 | 255 | 295 | 332 | 413 | 434 | 462 | 471 | 459 | 441 | 472 | 439 | 476 | 502 | 473 | 439 | 456 | 442 | 459 | 409 | 312 | 279 | 252 | 226 | 172 | 128 | 103 | 109 | 80 | 65 | 51 | 45 | 46 | 26 | 33 | 24 | 21 |
| tg | tg | 12683 | 42.1 | 49 | 7.5e-01 | 12 | -1.08 | 13 | 36 | 33 | 43 | 53 | 45 | 45 | 60 | 74 | 93 | 106 | 154 | 176 | 191 | 231 | 285 | 337 | 368 | 419 | 507 | 464 | 491 | 521 | 512 | 533 | 540 | 522 | 522 | 540 | 517 | 533 | 524 | 494 | 439 | 390 | 331 | 271 | 207 | 194 | 163 | 145 | 122 | 93 | 74 | 58 | 55 | 44 | 44 | 31 | 28 | 25 |
| ga | ga | 15201 | 39.6 | 49 | 8.3e-01 | 13 | -1.12 | 14 | 28 | 35 | 40 | 59 | 43 | 66 | 64 | 95 | 105 | 125 | 166 | 198 | 217 | 271 | 336 | 385 | 448 | 538 | 611 | 665 | 628 | 609 | 618 | 654 | 622 | 608 | 662 | 603 | 624 | 592 | 618 | 646 | 538 | 444 | 371 | 335 | 278 | 225 | 194 | 151 | 115 | 112 | 91 | 85 | 58 | 64 | 48 | 45 | 36 | 32 |
| gt | gt | 14025 | 35.1 | 49 | 9.3e-01 | 15 | -1.17 | 15 | 37 | 41 | 37 | 41 | 53 | 53 | 72 | 88 | 86 | 109 | 142 | 182 | 212 | 251 | 330 | 370 | 400 | 477 | 547 | 573 | 522 | 572 | 569 | 582 | 604 | 586 | 614 | 594 | 589 | 572 | 574 | 546 | 512 | 448 | 357 | 296 | 267 | 222 | 170 | 150 | 111 | 97 | 69 | 51 | 60 | 52 | 49 | 35 | 31 | 23 |
| cg | cg | 33859 | 28.5 | 49 | 9.9e-01 | 16 | -1.20 | 16 | 66 | 75 | 95 | 82 | 116 | 137 | 162 | 189 | 221 | 298 | 359 | 416 | 498 | 642 | 727 | 871 | 1062 | 1214 | 1315 | 1429 | 1381 | 1372 | 1361 | 1354 | 1391 | 1349 | 1371 | 1393 | 1399 | 1398 | 1408 | 1320 | 1209 | 1094 | 905 | 730 | 671 | 523 | 433 | 371 | 271 | 228 | 196 | 171 | 138 | 108 | 107 | 92 | 79 | 62 |
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