position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/01/22/peak-motifs.2026-01-22.084501_2026-01-22.084501_yybHyF/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title WT48h_sup3 -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/01/22/peak-motifs.2026-01-22.084501_2026-01-22.084501_yybHyF/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/01/22/peak-motifs.2026-01-22.084501_2026-01-22.084501_yybHyF/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/01/22/peak-motifs.2026-01-22.084501_2026-01-22.084501_yybHyF/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 493 Sum of sequence lengths 308249 Min sequence length 0 Max sequence length 1000 Average sequence length 625 Possible positions 307756 Oligonucleotide statistics: Total occurrences 303121 Position interval parameters: Position interval 20 Number of windows 50 Total positions 307756 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 75 1410 2 -24 [-479,-460] -469.5 84 1549 3 -23 [-459,-440] -449.5 96 1802 4 -22 [-439,-420] -429.5 109 2018 5 -21 [-419,-400] -409.5 119 2277 6 -20 [-399,-380] -389.5 139 2605 7 -19 [-379,-360] -369.5 165 3049 8 -18 [-359,-340] -349.5 179 3447 9 -17 [-339,-320] -329.5 205 3837 10 -16 [-319,-300] -309.5 233 4406 11 -15 [-299,-280] -289.5 262 5015 12 -14 [-279,-260] -269.5 297 5582 13 -13 [-259,-240] -249.5 330 6229 14 -12 [-239,-220] -229.5 368 7022 15 -11 [-219,-200] -209.5 397 7712 16 -10 [-199,-180] -189.5 442 8468 17 -9 [-179,-160] -169.5 475 9188 18 -8 [-159,-140] -149.5 490 9703 19 -7 [-139,-120] -129.5 493 9845 20 -6 [-119,-100] -109.5 493 9860 21 -5 [-99,-80] -89.5 493 9860 22 -4 [-79,-60] -69.5 493 9860 23 -3 [-59,-40] -49.5 493 9860 24 -2 [-39,-20] -29.5 493 9860 25 -1 [-19,0] -9.5 493 9860 26 0 [1,20] 10.5 493 9860 27 1 [21,40] 30.5 493 9860 28 2 [41,60] 50.5 493 9860 29 3 [61,80] 70.5 493 9860 30 4 [81,100] 90.5 493 9860 31 5 [101,120] 110.5 493 9860 32 6 [121,140] 130.5 493 9840 33 7 [141,160] 150.5 490 9671 34 8 [161,180] 170.5 471 9123 35 9 [181,200] 190.5 441 8386 36 10 [201,220] 210.5 397 7652 37 11 [221,240] 230.5 365 6953 38 12 [241,260] 250.5 328 6164 39 13 [261,280] 270.5 293 5517 40 14 [281,300] 290.5 260 4960 41 15 [301,320] 310.5 233 4350 42 16 [321,340] 330.5 204 3785 43 17 [341,360] 350.5 178 3423 44 18 [361,380] 370.5 165 2994 45 19 [381,400] 390.5 136 2565 46 20 [401,420] 410.5 119 2257 47 21 [421,440] 430.5 108 1996 48 22 [441,460] 450.5 96 1771 49 23 [461,480] 470.5 81 1535 50 24 [481,500] 490.5 75 1330 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| tt | tt | 5198 | 119.0 | 49 | 9.5e-08 | 1.5e-06 | 5.82 | 1 | 27 | 29 | 27 | 34 | 37 | 36 | 52 | 60 | 60 | 81 | 60 | 88 | 112 | 123 | 145 | 124 | 135 | 153 | 153 | 180 | 158 | 187 | 186 | 187 | 202 | 213 | 191 | 192 | 204 | 203 | 147 | 153 | 157 | 134 | 124 | 128 | 100 | 76 | 75 | 77 | 69 | 48 | 47 | 37 | 55 | 47 | 27 | 19 | 19 | 20 |
| aa | aa | 5238 | 91.3 | 49 | 2.3e-04 | 0.0037 | 2.43 | 2 | 25 | 28 | 33 | 41 | 37 | 47 | 66 | 57 | 60 | 52 | 79 | 104 | 105 | 109 | 132 | 124 | 131 | 144 | 170 | 183 | 174 | 195 | 218 | 198 | 192 | 191 | 191 | 156 | 180 | 142 | 163 | 187 | 154 | 132 | 127 | 107 | 114 | 102 | 93 | 68 | 65 | 69 | 64 | 53 | 46 | 34 | 21 | 26 | 27 | 22 |
| at | at | 5578 | 72.0 | 49 | 1.8e-02 | 0.28 | 0.55 | 3 | 24 | 36 | 43 | 44 | 53 | 43 | 52 | 52 | 90 | 91 | 85 | 103 | 103 | 121 | 144 | 163 | 176 | 182 | 162 | 187 | 183 | 185 | 174 | 197 | 190 | 160 | 141 | 181 | 183 | 191 | 191 | 220 | 159 | 155 | 142 | 128 | 86 | 107 | 106 | 88 | 69 | 66 | 57 | 53 | 43 | 40 | 42 | 29 | 27 | 31 |
| ct | ct | 15000 | 63.2 | 49 | 8.3e-02 | 1.3 | -0.12 | 4 | 74 | 67 | 67 | 100 | 89 | 129 | 126 | 156 | 203 | 219 | 202 | 258 | 340 | 358 | 360 | 413 | 444 | 464 | 496 | 473 | 496 | 466 | 465 | 468 | 506 | 488 | 449 | 478 | 489 | 466 | 469 | 465 | 485 | 451 | 430 | 400 | 335 | 298 | 262 | 258 | 250 | 205 | 158 | 135 | 135 | 119 | 94 | 76 | 80 | 86 |
| gg | gg | 33398 | 62.0 | 49 | 1.0e-01 | 1.6 | -0.21 | 5 | 140 | 162 | 210 | 218 | 267 | 244 | 362 | 343 | 412 | 480 | 563 | 590 | 664 | 802 | 927 | 913 | 1019 | 1033 | 1064 | 1031 | 1043 | 1060 | 1089 | 1033 | 1014 | 1026 | 1081 | 1049 | 1048 | 1090 | 1061 | 1091 | 1015 | 1059 | 918 | 846 | 778 | 687 | 621 | 577 | 463 | 423 | 341 | 348 | 287 | 232 | 221 | 180 | 152 | 121 |
| tc | tc | 19408 | 55.9 | 49 | 2.3e-01 | 3.7 | -0.57 | 6 | 88 | 105 | 113 | 114 | 138 | 163 | 189 | 229 | 270 | 284 | 263 | 342 | 421 | 456 | 469 | 543 | 569 | 615 | 612 | 658 | 615 | 587 | 618 | 637 | 647 | 566 | 581 | 597 | 641 | 609 | 626 | 620 | 620 | 616 | 551 | 504 | 431 | 402 | 362 | 346 | 270 | 237 | 196 | 168 | 151 | 165 | 130 | 92 | 96 | 86 |
| cc | cc | 33610 | 54.8 | 49 | 2.6e-01 | 4.2 | -0.63 | 7 | 154 | 163 | 183 | 214 | 233 | 304 | 316 | 389 | 410 | 483 | 556 | 595 | 707 | 740 | 792 | 933 | 1076 | 1073 | 1101 | 1098 | 1121 | 1076 | 1089 | 1052 | 1022 | 1138 | 1047 | 1066 | 1048 | 1053 | 1074 | 1076 | 1009 | 1003 | 982 | 924 | 759 | 655 | 583 | 501 | 470 | 383 | 373 | 332 | 274 | 247 | 211 | 218 | 164 | 140 |
| ac | ac | 18231 | 54.4 | 49 | 2.8e-01 | 4.4 | -0.65 | 8 | 69 | 89 | 99 | 109 | 129 | 164 | 181 | 208 | 229 | 236 | 292 | 347 | 378 | 402 | 426 | 506 | 537 | 581 | 592 | 578 | 548 | 609 | 589 | 573 | 587 | 588 | 612 | 560 | 575 | 576 | 585 | 552 | 598 | 522 | 463 | 454 | 432 | 393 | 342 | 275 | 255 | 229 | 249 | 191 | 145 | 141 | 105 | 126 | 105 | 100 |
| ca | ca | 15875 | 52.7 | 49 | 3.3e-01 | 5.3 | -0.73 | 9 | 82 | 91 | 86 | 103 | 137 | 154 | 164 | 184 | 198 | 226 | 257 | 293 | 345 | 354 | 397 | 466 | 467 | 512 | 472 | 466 | 488 | 535 | 517 | 529 | 487 | 496 | 527 | 514 | 492 | 512 | 510 | 498 | 495 | 435 | 415 | 363 | 380 | 337 | 276 | 223 | 233 | 186 | 197 | 149 | 144 | 117 | 87 | 107 | 90 | 82 |
| gc | gc | 37085 | 50.8 | 49 | 4.0e-01 | 6.5 | -0.81 | 10 | 191 | 162 | 212 | 248 | 305 | 310 | 359 | 418 | 432 | 530 | 640 | 664 | 718 | 843 | 908 | 1048 | 1130 | 1178 | 1159 | 1125 | 1200 | 1187 | 1136 | 1138 | 1154 | 1201 | 1158 | 1218 | 1142 | 1203 | 1193 | 1148 | 1160 | 1101 | 1082 | 924 | 886 | 736 | 667 | 595 | 569 | 476 | 426 | 392 | 332 | 264 | 256 | 222 | 184 | 155 |
| ta | ta | 3042 | 46.2 | 49 | 5.9e-01 | 9.4 | -0.97 | 11 | 15 | 15 | 16 | 19 | 21 | 27 | 29 | 26 | 34 | 51 | 49 | 60 | 60 | 56 | 94 | 79 | 81 | 97 | 102 | 108 | 100 | 94 | 93 | 103 | 122 | 91 | 87 | 109 | 97 | 92 | 98 | 98 | 100 | 85 | 73 | 78 | 57 | 69 | 40 | 41 | 47 | 47 | 32 | 31 | 23 | 16 | 21 | 17 | 24 | 18 |
| gt | gt | 17716 | 44.3 | 49 | 6.6e-01 | 11 | -1.02 | 12 | 92 | 107 | 119 | 118 | 124 | 144 | 192 | 211 | 225 | 267 | 277 | 329 | 348 | 390 | 466 | 473 | 476 | 548 | 578 | 559 | 535 | 531 | 572 | 583 | 589 | 531 | 589 | 559 | 585 | 539 | 568 | 557 | 573 | 547 | 488 | 437 | 408 | 368 | 327 | 318 | 245 | 217 | 187 | 160 | 136 | 135 | 125 | 99 | 101 | 64 |
| cg | cg | 43881 | 41.6 | 49 | 7.6e-01 | 12 | -1.09 | 13 | 200 | 201 | 264 | 270 | 335 | 382 | 431 | 506 | 529 | 626 | 725 | 788 | 841 | 1001 | 1051 | 1200 | 1325 | 1383 | 1416 | 1420 | 1374 | 1370 | 1383 | 1336 | 1395 | 1356 | 1392 | 1411 | 1365 | 1407 | 1405 | 1384 | 1386 | 1342 | 1260 | 1102 | 1047 | 901 | 827 | 755 | 611 | 546 | 517 | 459 | 358 | 324 | 313 | 266 | 216 | 179 |
| ag | ag | 14549 | 39.9 | 49 | 8.2e-01 | 13 | -1.12 | 14 | 67 | 77 | 78 | 99 | 118 | 143 | 148 | 173 | 166 | 205 | 269 | 257 | 290 | 338 | 382 | 402 | 414 | 466 | 434 | 477 | 489 | 473 | 453 | 496 | 426 | 470 | 491 | 457 | 451 | 470 | 445 | 468 | 465 | 396 | 368 | 349 | 352 | 296 | 243 | 219 | 218 | 197 | 165 | 137 | 131 | 110 | 99 | 79 | 70 | 63 |
| tg | tg | 15853 | 36.0 | 49 | 9.2e-01 | 15 | -1.17 | 15 | 84 | 93 | 102 | 125 | 111 | 118 | 153 | 175 | 219 | 223 | 267 | 288 | 316 | 357 | 396 | 422 | 461 | 485 | 507 | 451 | 502 | 499 | 494 | 510 | 523 | 522 | 506 | 514 | 519 | 489 | 521 | 504 | 496 | 473 | 425 | 391 | 336 | 306 | 294 | 279 | 245 | 201 | 177 | 151 | 142 | 113 | 107 | 97 | 95 | 69 |
| ga | ga | 19459 | 33.0 | 49 | 9.6e-01 | 15 | -1.19 | 16 | 71 | 97 | 110 | 139 | 134 | 176 | 189 | 220 | 250 | 263 | 331 | 364 | 357 | 449 | 480 | 519 | 577 | 608 | 627 | 666 | 642 | 626 | 604 | 632 | 604 | 623 | 624 | 588 | 624 | 628 | 612 | 638 | 622 | 562 | 494 | 481 | 435 | 399 | 368 | 320 | 266 | 255 | 237 | 198 | 163 | 153 | 137 | 118 | 85 | 94 |
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