position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/01/25/peak-motifs.2026-01-25.180830_2026-01-25.180830_qbHDLg/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title Merged -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/01/25/peak-motifs.2026-01-25.180830_2026-01-25.180830_qbHDLg/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/01/25/peak-motifs.2026-01-25.180830_2026-01-25.180830_qbHDLg/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/01/25/peak-motifs.2026-01-25.180830_2026-01-25.180830_qbHDLg/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 5196 Sum of sequence lengths 3095499 Min sequence length 0 Max sequence length 1000 Average sequence length 595 Possible positions 3090303 Oligonucleotide statistics: Total occurrences 3086572 Position interval parameters: Position interval 20 Number of windows 50 Total positions 3090303 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 910 17618 2 -24 [-479,-460] -469.5 994 19100 3 -23 [-459,-440] -449.5 1075 20743 4 -22 [-439,-420] -429.5 1166 22523 5 -21 [-419,-400] -409.5 1286 24612 6 -20 [-399,-380] -389.5 1396 26774 7 -19 [-379,-360] -369.5 1547 29536 8 -18 [-359,-340] -349.5 1688 32474 9 -17 [-339,-320] -329.5 1861 35427 10 -16 [-319,-300] -309.5 2076 39345 11 -15 [-299,-280] -289.5 2280 43683 12 -14 [-279,-260] -269.5 2558 48607 13 -13 [-259,-240] -249.5 2873 54481 14 -12 [-239,-220] -229.5 3256 61297 15 -11 [-219,-200] -209.5 3730 70170 16 -10 [-199,-180] -189.5 4304 80606 17 -9 [-179,-160] -169.5 4922 92773 18 -8 [-159,-140] -149.5 5196 102732 19 -7 [-139,-120] -129.5 5196 103920 20 -6 [-119,-100] -109.5 5196 103920 21 -5 [-99,-80] -89.5 5196 103920 22 -4 [-79,-60] -69.5 5196 103920 23 -3 [-59,-40] -49.5 5196 103920 24 -2 [-39,-20] -29.5 5196 103920 25 -1 [-19,0] -9.5 5196 103920 26 0 [1,20] 10.5 5196 103920 27 1 [21,40] 30.5 5196 103920 28 2 [41,60] 50.5 5196 103920 29 3 [61,80] 70.5 5196 103920 30 4 [81,100] 90.5 5196 103920 31 5 [101,120] 110.5 5196 103920 32 6 [121,140] 130.5 5196 103920 33 7 [141,160] 150.5 5196 102156 34 8 [161,180] 170.5 4873 91561 35 9 [181,200] 190.5 4251 79461 36 10 [201,220] 210.5 3694 69205 37 11 [221,240] 230.5 3225 60513 38 12 [241,260] 250.5 2852 53857 39 13 [261,280] 270.5 2536 48048 40 14 [281,300] 290.5 2268 43278 41 15 [301,320] 310.5 2054 38939 42 16 [321,340] 330.5 1843 35081 43 17 [341,360] 350.5 1677 32187 44 18 [361,380] 370.5 1532 29226 45 19 [381,400] 390.5 1384 26542 46 20 [401,420] 410.5 1278 24370 47 21 [421,440] 430.5 1158 22339 48 22 [441,460] 450.5 1071 20571 49 23 [461,480] 470.5 985 18935 50 24 [481,500] 490.5 903 16653 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| aa | aa | 256561 | 357.4 | 49 | 1.9e-48 | 3.1e-47 | 46.51 | 1 | 1552 | 1717 | 1850 | 2017 | 2088 | 2245 | 2581 | 2733 | 3017 | 3373 | 3716 | 4002 | 4796 | 5195 | 5935 | 6786 | 7683 | 8424 | 8301 | 8496 | 8435 | 8377 | 8356 | 8262 | 8238 | 8353 | 8488 | 8234 | 8326 | 8463 | 8510 | 8398 | 8282 | 7617 | 6811 | 5761 | 5254 | 4560 | 4102 | 3726 | 3285 | 3165 | 2916 | 2559 | 2294 | 2293 | 2048 | 1772 | 1678 | 1491 |
| tt | tt | 255606 | 263.0 | 49 | 4.7e-31 | 7.5e-30 | 29.13 | 2 | 1469 | 1655 | 1822 | 1963 | 2205 | 2376 | 2537 | 2859 | 3098 | 3469 | 3630 | 4047 | 4522 | 5046 | 5968 | 6675 | 7691 | 8437 | 8494 | 8194 | 8626 | 8471 | 8271 | 8240 | 8384 | 8199 | 8193 | 8354 | 8472 | 8355 | 8566 | 8531 | 8444 | 7607 | 6677 | 5860 | 5004 | 4526 | 4200 | 3646 | 3354 | 2929 | 2773 | 2487 | 2314 | 2042 | 2009 | 1764 | 1683 | 1468 |
| ta | ta | 148425 | 244.6 | 49 | 8.5e-28 | 1.4e-26 | 25.86 | 3 | 904 | 983 | 1050 | 1112 | 1279 | 1386 | 1451 | 1656 | 1804 | 1985 | 2163 | 2393 | 2681 | 3062 | 3317 | 3907 | 4459 | 4887 | 5096 | 4921 | 4933 | 4725 | 4697 | 4781 | 4830 | 4794 | 4750 | 4782 | 4755 | 4829 | 4849 | 4770 | 4857 | 4484 | 3950 | 3451 | 3036 | 2763 | 2345 | 2172 | 1945 | 1764 | 1628 | 1489 | 1333 | 1189 | 1198 | 1059 | 901 | 870 |
| cc | cc | 166692 | 218.4 | 49 | 3.0e-23 | 4.8e-22 | 21.32 | 4 | 951 | 882 | 998 | 1171 | 1227 | 1428 | 1480 | 1725 | 1757 | 2031 | 2309 | 2544 | 2775 | 3155 | 3724 | 4319 | 4906 | 5596 | 5695 | 5638 | 5766 | 5655 | 5911 | 5856 | 5828 | 5749 | 5694 | 5697 | 5872 | 5687 | 5930 | 5780 | 5430 | 4903 | 4206 | 3688 | 3248 | 2768 | 2523 | 2358 | 2086 | 1838 | 1707 | 1603 | 1361 | 1271 | 1089 | 1055 | 954 | 868 |
| gc | gc | 218704 | 194.3 | 49 | 3.4e-19 | 5.4e-18 | 17.27 | 5 | 1135 | 1280 | 1420 | 1529 | 1634 | 1796 | 2052 | 2232 | 2479 | 2772 | 3042 | 3408 | 3797 | 4316 | 4914 | 5581 | 6432 | 7298 | 7245 | 7501 | 7384 | 7581 | 7607 | 7674 | 7535 | 7737 | 7692 | 7670 | 7483 | 7474 | 7384 | 7353 | 7277 | 6461 | 5427 | 4820 | 4287 | 3842 | 3303 | 2917 | 2682 | 2368 | 2202 | 1940 | 1818 | 1678 | 1430 | 1369 | 1317 | 1129 |
| cg | cg | 175611 | 155.2 | 49 | 5.8e-13 | 9.2e-12 | 11.03 | 6 | 920 | 1030 | 1157 | 1224 | 1331 | 1490 | 1614 | 1849 | 1990 | 2209 | 2441 | 2739 | 3069 | 3550 | 3881 | 4544 | 5062 | 5925 | 5803 | 6133 | 6000 | 6083 | 6051 | 6074 | 6129 | 6176 | 6173 | 6174 | 5969 | 5942 | 5960 | 5933 | 5793 | 5089 | 4424 | 3886 | 3342 | 3020 | 2611 | 2351 | 2186 | 1939 | 1778 | 1562 | 1491 | 1307 | 1229 | 1112 | 983 | 883 |
| at | at | 200346 | 142.8 | 49 | 4.2e-11 | 6.8e-10 | 9.17 | 7 | 1171 | 1301 | 1385 | 1555 | 1660 | 1789 | 1957 | 2121 | 2376 | 2604 | 2831 | 3303 | 3554 | 4045 | 4556 | 5294 | 6091 | 6570 | 6688 | 6714 | 6743 | 6508 | 6500 | 6610 | 6626 | 6552 | 6543 | 6456 | 6547 | 6580 | 6564 | 6676 | 6542 | 6038 | 5344 | 4602 | 4044 | 3607 | 3129 | 2900 | 2606 | 2410 | 2146 | 1889 | 1734 | 1586 | 1583 | 1396 | 1205 | 1115 |
| gg | gg | 165072 | 118.7 | 49 | 1.0e-07 | 1.7e-06 | 5.78 | 8 | 946 | 958 | 1114 | 1121 | 1284 | 1432 | 1581 | 1708 | 1855 | 2023 | 2364 | 2528 | 2865 | 3316 | 3823 | 4256 | 4923 | 5485 | 5608 | 5687 | 5648 | 5655 | 5576 | 5661 | 5711 | 5636 | 5605 | 5700 | 5620 | 5635 | 5715 | 5576 | 5487 | 4810 | 4257 | 3727 | 3187 | 2805 | 2592 | 2191 | 2002 | 1901 | 1626 | 1501 | 1322 | 1253 | 1048 | 1050 | 904 | 794 |
| ct | ct | 177680 | 76.9 | 49 | 6.6e-03 | 0.11 | 0.97 | 9 | 1039 | 1107 | 1166 | 1256 | 1412 | 1538 | 1646 | 1860 | 2109 | 2195 | 2534 | 2889 | 3147 | 3468 | 4007 | 4726 | 5402 | 5990 | 6062 | 6015 | 6097 | 5973 | 6063 | 5986 | 5900 | 5970 | 6050 | 6070 | 6142 | 5941 | 5970 | 5979 | 5934 | 5305 | 4355 | 4006 | 3445 | 3086 | 2715 | 2387 | 2136 | 1878 | 1754 | 1685 | 1540 | 1340 | 1213 | 1121 | 1110 | 961 |
| tc | tc | 182553 | 69.8 | 49 | 2.7e-02 | 0.44 | 0.36 | 10 | 1067 | 1128 | 1188 | 1325 | 1480 | 1602 | 1684 | 1912 | 2094 | 2286 | 2553 | 2925 | 3190 | 3545 | 4061 | 4874 | 5463 | 6185 | 6260 | 6201 | 6232 | 6154 | 6219 | 6224 | 6127 | 6117 | 6184 | 6314 | 6362 | 6178 | 6097 | 6232 | 6094 | 5357 | 4584 | 4030 | 3419 | 3043 | 2733 | 2518 | 2283 | 2013 | 1824 | 1708 | 1553 | 1365 | 1305 | 1195 | 1085 | 981 |
| ca | ca | 218952 | 63.2 | 49 | 8.3e-02 | 1.3 | -0.12 | 11 | 1283 | 1348 | 1440 | 1630 | 1738 | 1840 | 2150 | 2264 | 2460 | 2810 | 3076 | 3325 | 3758 | 4204 | 5023 | 5770 | 6645 | 7315 | 7441 | 7368 | 7168 | 7461 | 7511 | 7489 | 7395 | 7428 | 7416 | 7452 | 7339 | 7610 | 7386 | 7367 | 7090 | 6405 | 5570 | 4721 | 4230 | 3764 | 3470 | 3031 | 2773 | 2500 | 2276 | 2059 | 1843 | 1754 | 1559 | 1479 | 1380 | 1138 |
| ac | ac | 170814 | 54.8 | 49 | 2.6e-01 | 4.2 | -0.63 | 12 | 1001 | 1091 | 1167 | 1233 | 1398 | 1465 | 1668 | 1848 | 1935 | 2164 | 2491 | 2594 | 3027 | 3310 | 3908 | 4567 | 5212 | 5732 | 5831 | 5800 | 5686 | 5727 | 5821 | 5682 | 5758 | 5740 | 5718 | 5703 | 5610 | 5829 | 5841 | 5683 | 5462 | 4957 | 4351 | 3691 | 3343 | 2977 | 2703 | 2331 | 2177 | 1926 | 1780 | 1669 | 1530 | 1305 | 1240 | 1185 | 1055 | 892 |
| ag | ag | 178590 | 52.0 | 49 | 3.6e-01 | 5.7 | -0.76 | 13 | 1016 | 1086 | 1163 | 1264 | 1357 | 1481 | 1734 | 1791 | 2030 | 2232 | 2473 | 2816 | 3128 | 3568 | 4073 | 4583 | 5414 | 5899 | 6117 | 5925 | 5903 | 6064 | 6009 | 6055 | 6079 | 6117 | 6006 | 6133 | 6072 | 6187 | 5905 | 6019 | 6060 | 5293 | 4571 | 3965 | 3411 | 3116 | 2827 | 2523 | 2220 | 2000 | 1815 | 1667 | 1510 | 1434 | 1226 | 1169 | 1129 | 955 |
| ga | ga | 181783 | 44.9 | 49 | 6.4e-01 | 10 | -1.01 | 14 | 1020 | 1121 | 1227 | 1286 | 1368 | 1538 | 1722 | 1871 | 2049 | 2189 | 2533 | 2948 | 3250 | 3688 | 4164 | 4728 | 5491 | 6069 | 6091 | 6163 | 6169 | 6178 | 6066 | 6120 | 6196 | 6188 | 6092 | 6094 | 6132 | 6190 | 6035 | 6231 | 6117 | 5338 | 4688 | 4100 | 3499 | 3144 | 2817 | 2600 | 2232 | 2066 | 1838 | 1658 | 1568 | 1404 | 1293 | 1193 | 1096 | 925 |
| tg | tg | 217869 | 39.1 | 49 | 8.4e-01 | 13 | -1.13 | 15 | 1177 | 1361 | 1455 | 1594 | 1729 | 1855 | 2059 | 2245 | 2450 | 2760 | 3063 | 3491 | 3878 | 4333 | 4939 | 5580 | 6661 | 7157 | 7299 | 7300 | 7333 | 7393 | 7356 | 7353 | 7311 | 7316 | 7436 | 7222 | 7300 | 7158 | 7309 | 7441 | 7328 | 6545 | 5678 | 4943 | 4335 | 3823 | 3324 | 3080 | 2704 | 2399 | 2272 | 2010 | 1829 | 1716 | 1543 | 1448 | 1386 | 1192 |
| gt | gt | 171314 | 30.9 | 49 | 9.8e-01 | 16 | -1.20 | 16 | 967 | 1052 | 1141 | 1243 | 1422 | 1513 | 1620 | 1800 | 1924 | 2223 | 2444 | 2647 | 3005 | 3442 | 3834 | 4372 | 5162 | 5659 | 5808 | 5764 | 5663 | 5734 | 5722 | 5726 | 5753 | 5730 | 5793 | 5747 | 5774 | 5684 | 5722 | 5780 | 5750 | 5150 | 4503 | 3878 | 3336 | 2947 | 2589 | 2472 | 2191 | 1925 | 1792 | 1647 | 1433 | 1369 | 1266 | 1175 | 1049 | 972 |
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