position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/01/25/peak-motifs.2026-01-25.181000_2026-01-25.181000_CpuIe4/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title S1 -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/01/25/peak-motifs.2026-01-25.181000_2026-01-25.181000_CpuIe4/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/01/25/peak-motifs.2026-01-25.181000_2026-01-25.181000_CpuIe4/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/01/25/peak-motifs.2026-01-25.181000_2026-01-25.181000_CpuIe4/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 2616 Sum of sequence lengths 1315716 Min sequence length 0 Max sequence length 1000 Average sequence length 502 Possible positions 1313100 Oligonucleotide statistics: Total occurrences 1305428 Position interval parameters: Position interval 20 Number of windows 50 Total positions 1313100 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 153 2818 2 -24 [-479,-460] -469.5 174 3331 3 -23 [-459,-440] -449.5 202 3789 4 -22 [-439,-420] -429.5 236 4438 5 -21 [-419,-400] -409.5 285 5253 6 -20 [-399,-380] -389.5 325 6126 7 -19 [-379,-360] -369.5 391 7230 8 -18 [-359,-340] -349.5 456 8634 9 -17 [-339,-320] -329.5 530 9793 10 -16 [-319,-300] -309.5 634 11628 11 -15 [-299,-280] -289.5 732 13727 12 -14 [-279,-260] -269.5 878 16087 13 -13 [-259,-240] -249.5 1034 19113 14 -12 [-239,-220] -229.5 1253 22771 15 -11 [-219,-200] -209.5 1568 28466 16 -10 [-199,-180] -189.5 1948 35350 17 -9 [-179,-160] -169.5 2352 43358 18 -8 [-159,-140] -149.5 2616 50950 19 -7 [-139,-120] -129.5 2616 52320 20 -6 [-119,-100] -109.5 2616 52320 21 -5 [-99,-80] -89.5 2616 52320 22 -4 [-79,-60] -69.5 2616 52320 23 -3 [-59,-40] -49.5 2616 52320 24 -2 [-39,-20] -29.5 2616 52320 25 -1 [-19,0] -9.5 2616 52320 26 0 [1,20] 10.5 2616 52320 27 1 [21,40] 30.5 2616 52320 28 2 [41,60] 50.5 2616 52320 29 3 [61,80] 70.5 2616 52320 30 4 [81,100] 90.5 2616 52320 31 5 [101,120] 110.5 2616 52320 32 6 [121,140] 130.5 2616 52320 33 7 [141,160] 150.5 2616 50398 34 8 [161,180] 170.5 2315 42569 35 9 [181,200] 190.5 1919 34612 36 10 [201,220] 210.5 1547 27828 37 11 [221,240] 230.5 1233 22324 38 12 [241,260] 250.5 1022 18804 39 13 [261,280] 270.5 862 15798 40 14 [281,300] 290.5 726 13536 41 15 [301,320] 310.5 627 11420 42 16 [321,340] 330.5 519 9643 43 17 [341,360] 350.5 453 8500 44 18 [361,380] 370.5 387 7097 45 19 [381,400] 390.5 324 6039 46 20 [401,420] 410.5 281 5158 47 21 [421,440] 430.5 233 4364 48 22 [441,460] 450.5 199 3729 49 23 [461,480] 470.5 173 3297 50 24 [481,500] 490.5 151 2642 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| aa | aa | 91956 | 161.1 | 49 | 7.2e-14 | 1.1e-12 | 11.94 | 1 | 196 | 242 | 267 | 342 | 338 | 371 | 494 | 626 | 647 | 794 | 954 | 1096 | 1435 | 1694 | 2064 | 2571 | 3084 | 3789 | 3761 | 3809 | 3624 | 3571 | 3513 | 3410 | 3462 | 3552 | 3467 | 3528 | 3710 | 3807 | 3752 | 3697 | 3600 | 3089 | 2582 | 1948 | 1589 | 1350 | 1078 | 911 | 764 | 644 | 559 | 500 | 398 | 355 | 293 | 216 | 219 | 194 |
| ta | ta | 53100 | 146.4 | 49 | 1.2e-11 | 2e-10 | 9.70 | 2 | 125 | 148 | 146 | 169 | 205 | 268 | 281 | 340 | 387 | 467 | 536 | 613 | 786 | 954 | 1165 | 1542 | 1827 | 2151 | 2258 | 2234 | 2118 | 2018 | 2009 | 1954 | 1974 | 1987 | 2026 | 2056 | 2018 | 2076 | 2104 | 2114 | 2138 | 1809 | 1522 | 1231 | 931 | 803 | 619 | 528 | 444 | 380 | 336 | 318 | 220 | 210 | 205 | 132 | 108 | 110 |
| gc | gc | 103098 | 136.0 | 49 | 4.1e-10 | 6.5e-09 | 8.19 | 3 | 180 | 238 | 274 | 350 | 417 | 495 | 570 | 674 | 791 | 927 | 1085 | 1314 | 1480 | 1758 | 2207 | 2676 | 3289 | 3827 | 3997 | 4082 | 4148 | 4210 | 4245 | 4297 | 4256 | 4376 | 4419 | 4278 | 4097 | 4104 | 3987 | 4026 | 3808 | 3340 | 2571 | 2098 | 1765 | 1416 | 1190 | 1049 | 872 | 738 | 653 | 532 | 494 | 397 | 347 | 281 | 272 | 201 |
| cg | cg | 81228 | 130.8 | 49 | 2.3e-09 | 3.6e-08 | 7.44 | 4 | 156 | 201 | 248 | 264 | 326 | 377 | 438 | 542 | 605 | 719 | 876 | 1027 | 1155 | 1390 | 1695 | 2083 | 2494 | 3095 | 3135 | 3250 | 3233 | 3299 | 3282 | 3429 | 3414 | 3460 | 3446 | 3400 | 3346 | 3226 | 3136 | 3154 | 2967 | 2619 | 2066 | 1640 | 1315 | 1156 | 912 | 815 | 729 | 574 | 525 | 420 | 405 | 320 | 281 | 239 | 181 | 163 |
| tt | tt | 91786 | 121.2 | 49 | 4.8e-08 | 7.6e-07 | 6.12 | 5 | 211 | 248 | 303 | 275 | 382 | 426 | 501 | 572 | 636 | 829 | 879 | 1066 | 1266 | 1517 | 2024 | 2554 | 3195 | 3617 | 3669 | 3629 | 3689 | 3658 | 3634 | 3612 | 3488 | 3511 | 3535 | 3559 | 3620 | 3598 | 3710 | 3681 | 3718 | 3126 | 2561 | 2063 | 1555 | 1312 | 1142 | 974 | 765 | 644 | 583 | 472 | 407 | 343 | 289 | 299 | 238 | 201 |
| cc | cc | 78731 | 119.9 | 49 | 7.1e-08 | 1.1e-06 | 5.95 | 6 | 175 | 203 | 237 | 259 | 323 | 399 | 439 | 526 | 567 | 694 | 793 | 981 | 1082 | 1329 | 1667 | 2126 | 2494 | 3007 | 3101 | 3063 | 3155 | 3192 | 3399 | 3286 | 3300 | 3271 | 3252 | 3221 | 3185 | 3136 | 3181 | 3022 | 2827 | 2442 | 1982 | 1584 | 1287 | 1084 | 944 | 862 | 710 | 554 | 524 | 424 | 337 | 286 | 233 | 238 | 207 | 141 |
| at | at | 77204 | 114.3 | 49 | 3.9e-07 | 6.2e-06 | 5.21 | 7 | 173 | 200 | 227 | 245 | 283 | 370 | 440 | 485 | 524 | 674 | 820 | 985 | 1157 | 1350 | 1714 | 2216 | 2643 | 3060 | 3222 | 3096 | 3002 | 2939 | 2992 | 2968 | 2910 | 2993 | 2932 | 2927 | 3027 | 3027 | 3082 | 3157 | 3050 | 2523 | 2161 | 1789 | 1321 | 1076 | 925 | 783 | 702 | 578 | 513 | 422 | 366 | 314 | 295 | 202 | 184 | 160 |
| ac | ac | 71700 | 101.6 | 49 | 1.5e-05 | 0.00024 | 3.62 | 8 | 164 | 192 | 248 | 285 | 325 | 353 | 375 | 487 | 572 | 701 | 803 | 865 | 1048 | 1253 | 1546 | 1949 | 2383 | 2956 | 2877 | 2893 | 2802 | 2911 | 2930 | 2840 | 2848 | 2793 | 2898 | 2837 | 2804 | 2896 | 2880 | 2693 | 2568 | 2290 | 1844 | 1411 | 1249 | 1043 | 894 | 723 | 628 | 496 | 444 | 379 | 323 | 268 | 251 | 190 | 174 | 118 |
| gg | gg | 78004 | 93.8 | 49 | 1.2e-04 | 0.002 | 2.70 | 9 | 158 | 194 | 239 | 266 | 311 | 365 | 469 | 544 | 640 | 660 | 817 | 957 | 1115 | 1373 | 1736 | 1930 | 2539 | 2894 | 3036 | 3083 | 3139 | 3135 | 2938 | 3181 | 3117 | 3092 | 3117 | 3179 | 3197 | 3116 | 3214 | 3193 | 2972 | 2428 | 2023 | 1665 | 1336 | 1103 | 965 | 776 | 651 | 653 | 545 | 404 | 386 | 320 | 254 | 256 | 175 | 148 |
| ca | ca | 94925 | 66.0 | 49 | 5.3e-02 | 0.85 | 0.07 | 10 | 219 | 246 | 268 | 353 | 402 | 465 | 544 | 589 | 712 | 923 | 1019 | 1136 | 1365 | 1647 | 2063 | 2582 | 3186 | 3751 | 3837 | 3775 | 3714 | 3807 | 3863 | 3867 | 3747 | 3815 | 3825 | 3778 | 3768 | 3887 | 3774 | 3669 | 3508 | 3029 | 2411 | 1967 | 1617 | 1321 | 1180 | 986 | 813 | 674 | 588 | 486 | 444 | 352 | 301 | 241 | 259 | 152 |
| gt | gt | 71842 | 60.6 | 49 | 1.2e-01 | 2 | -0.30 | 11 | 160 | 180 | 166 | 229 | 286 | 317 | 381 | 467 | 533 | 602 | 732 | 798 | 1013 | 1200 | 1497 | 1910 | 2379 | 2785 | 2826 | 2858 | 2910 | 2847 | 2791 | 2831 | 2911 | 2864 | 2860 | 2834 | 2907 | 2865 | 2900 | 2989 | 2871 | 2425 | 1913 | 1541 | 1233 | 1072 | 832 | 754 | 595 | 532 | 464 | 403 | 311 | 308 | 231 | 224 | 158 | 147 |
| ga | ga | 79975 | 57.4 | 49 | 1.9e-01 | 3.1 | -0.49 | 12 | 176 | 177 | 251 | 276 | 297 | 355 | 460 | 542 | 644 | 714 | 885 | 1038 | 1241 | 1443 | 1757 | 2125 | 2610 | 3121 | 3204 | 3124 | 3148 | 3214 | 3051 | 3172 | 3238 | 3168 | 3103 | 3115 | 3226 | 3170 | 3146 | 3229 | 3103 | 2495 | 2103 | 1679 | 1303 | 1149 | 986 | 826 | 721 | 632 | 544 | 436 | 386 | 312 | 275 | 241 | 197 | 167 |
| ct | ct | 78403 | 56.8 | 49 | 2.1e-01 | 3.3 | -0.52 | 13 | 168 | 207 | 229 | 272 | 331 | 384 | 397 | 541 | 618 | 676 | 837 | 1028 | 1157 | 1357 | 1691 | 2121 | 2610 | 2997 | 3068 | 3145 | 3161 | 3117 | 3278 | 3043 | 3155 | 3059 | 3170 | 3211 | 3070 | 3083 | 3095 | 3101 | 2966 | 2446 | 2008 | 1656 | 1379 | 1134 | 957 | 786 | 652 | 539 | 500 | 421 | 353 | 320 | 268 | 225 | 232 | 184 |
| tg | tg | 94466 | 54.1 | 49 | 2.9e-01 | 4.6 | -0.66 | 14 | 176 | 236 | 245 | 328 | 364 | 437 | 510 | 656 | 690 | 802 | 984 | 1189 | 1383 | 1567 | 2051 | 2511 | 3198 | 3619 | 3683 | 3661 | 3815 | 3785 | 3740 | 3750 | 3778 | 3753 | 3803 | 3696 | 3699 | 3647 | 3871 | 3937 | 3683 | 3052 | 2512 | 2018 | 1652 | 1329 | 1114 | 997 | 797 | 705 | 619 | 537 | 418 | 397 | 327 | 284 | 254 | 207 |
| ag | ag | 79365 | 45.1 | 49 | 6.3e-01 | 10 | -1.01 | 15 | 181 | 173 | 205 | 260 | 307 | 357 | 477 | 506 | 638 | 738 | 823 | 962 | 1191 | 1425 | 1739 | 2104 | 2619 | 3036 | 3190 | 3160 | 3182 | 3175 | 3083 | 3141 | 3232 | 3159 | 3104 | 3190 | 3144 | 3238 | 3058 | 3155 | 3153 | 2585 | 2052 | 1643 | 1301 | 1174 | 968 | 809 | 695 | 606 | 500 | 442 | 364 | 294 | 253 | 225 | 191 | 158 |
| tc | tc | 79645 | 40.4 | 49 | 8.1e-01 | 13 | -1.11 | 16 | 180 | 226 | 216 | 245 | 340 | 367 | 434 | 517 | 569 | 684 | 850 | 1009 | 1179 | 1377 | 1658 | 2165 | 2602 | 3035 | 3213 | 3161 | 3161 | 3129 | 3251 | 3231 | 3168 | 3182 | 3138 | 3248 | 3250 | 3198 | 3159 | 3186 | 3051 | 2482 | 2052 | 1707 | 1322 | 1137 | 947 | 810 | 724 | 562 | 495 | 429 | 387 | 325 | 256 | 231 | 228 | 172 |
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