position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/01/25/peak-motifs.2026-01-25.181029_2026-01-25.181029_PUJ88V/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title S2 -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/01/25/peak-motifs.2026-01-25.181029_2026-01-25.181029_PUJ88V/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/01/25/peak-motifs.2026-01-25.181029_2026-01-25.181029_PUJ88V/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/01/25/peak-motifs.2026-01-25.181029_2026-01-25.181029_PUJ88V/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 3920 Sum of sequence lengths 2594017 Min sequence length 0 Max sequence length 1000 Average sequence length 661 Possible positions 2590097 Oligonucleotide statistics: Total occurrences 2582553 Position interval parameters: Position interval 20 Number of windows 50 Total positions 2590097 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 1007 19615 2 -24 [-479,-460] -469.5 1090 21013 3 -23 [-459,-440] -449.5 1171 22646 4 -22 [-439,-420] -429.5 1256 24361 5 -21 [-419,-400] -409.5 1354 26175 6 -20 [-399,-380] -389.5 1456 28089 7 -19 [-379,-360] -369.5 1573 30369 8 -18 [-359,-340] -349.5 1690 32659 9 -17 [-339,-320] -329.5 1835 35290 10 -16 [-319,-300] -309.5 1995 38326 11 -15 [-299,-280] -289.5 2159 41644 12 -14 [-279,-260] -269.5 2378 45608 13 -13 [-259,-240] -249.5 2606 50103 14 -12 [-239,-220] -229.5 2862 54771 15 -11 [-219,-200] -209.5 3153 60309 16 -10 [-199,-180] -189.5 3489 66709 17 -9 [-179,-160] -169.5 3856 73672 18 -8 [-159,-140] -149.5 3920 78347 19 -7 [-139,-120] -129.5 3920 78400 20 -6 [-119,-100] -109.5 3920 78400 21 -5 [-99,-80] -89.5 3920 78400 22 -4 [-79,-60] -69.5 3920 78400 23 -3 [-59,-40] -49.5 3920 78400 24 -2 [-39,-20] -29.5 3920 78400 25 -1 [-19,0] -9.5 3920 78400 26 0 [1,20] 10.5 3920 78400 27 1 [21,40] 30.5 3920 78400 28 2 [41,60] 50.5 3920 78400 29 3 [61,80] 70.5 3920 78400 30 4 [81,100] 90.5 3920 78400 31 5 [101,120] 110.5 3920 78400 32 6 [121,140] 130.5 3920 78400 33 7 [141,160] 150.5 3920 78203 34 8 [161,180] 170.5 3832 72951 35 9 [181,200] 190.5 3460 66033 36 10 [201,220] 210.5 3130 59718 37 11 [221,240] 230.5 2842 54251 38 12 [241,260] 250.5 2592 49640 39 13 [261,280] 270.5 2364 45171 40 14 [281,300] 290.5 2147 41320 41 15 [301,320] 310.5 1979 38029 42 16 [321,340] 330.5 1824 34999 43 17 [341,360] 350.5 1679 32425 44 18 [361,380] 370.5 1562 30128 45 19 [381,400] 390.5 1444 27876 46 20 [401,420] 410.5 1346 25979 47 21 [421,440] 430.5 1248 24198 48 22 [441,460] 450.5 1166 22471 49 23 [461,480] 470.5 1079 20843 50 24 [481,500] 490.5 1000 18556 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| aa | aa | 228195 | 116.3 | 49 | 2.2e-07 | 3.5e-06 | 5.46 | 1 | 1736 | 1902 | 2014 | 2179 | 2292 | 2449 | 2753 | 2849 | 3089 | 3388 | 3711 | 3996 | 4514 | 4843 | 5359 | 5922 | 6511 | 6725 | 6728 | 6873 | 6813 | 6866 | 6757 | 6895 | 6831 | 6867 | 7035 | 6646 | 6664 | 6738 | 6880 | 6863 | 6768 | 6414 | 5907 | 5262 | 4863 | 4321 | 4009 | 3735 | 3316 | 3211 | 3052 | 2710 | 2522 | 2507 | 2269 | 2042 | 1919 | 1680 |
| cc | cc | 131951 | 107.5 | 49 | 2.9e-06 | 4.6e-05 | 4.34 | 2 | 1050 | 958 | 1084 | 1248 | 1289 | 1435 | 1504 | 1666 | 1729 | 1905 | 2081 | 2198 | 2472 | 2650 | 3057 | 3333 | 3670 | 4079 | 4075 | 4013 | 4095 | 3965 | 4015 | 4008 | 4055 | 3868 | 3942 | 3931 | 4167 | 4010 | 4205 | 4201 | 4045 | 3794 | 3351 | 3048 | 2760 | 2465 | 2290 | 2128 | 1918 | 1784 | 1729 | 1627 | 1427 | 1358 | 1170 | 1119 | 1031 | 949 |
| tt | tt | 228313 | 94.4 | 49 | 1.1e-04 | 0.0017 | 2.77 | 3 | 1587 | 1787 | 2006 | 2204 | 2379 | 2587 | 2702 | 2957 | 3235 | 3512 | 3664 | 3976 | 4525 | 4840 | 5391 | 5838 | 6467 | 6972 | 6915 | 6593 | 7063 | 6898 | 6825 | 6682 | 6927 | 6731 | 6811 | 6903 | 7004 | 6919 | 6993 | 7004 | 6880 | 6350 | 5804 | 5278 | 4632 | 4375 | 4068 | 3591 | 3360 | 3096 | 2846 | 2667 | 2513 | 2219 | 2214 | 1965 | 1893 | 1665 |
| gg | gg | 131000 | 62.1 | 49 | 9.9e-02 | 1.6 | -0.20 | 4 | 1046 | 1048 | 1169 | 1223 | 1349 | 1478 | 1531 | 1683 | 1759 | 1915 | 2147 | 2288 | 2541 | 2841 | 3116 | 3448 | 3747 | 3955 | 3979 | 4027 | 3893 | 3921 | 3975 | 3913 | 4068 | 4028 | 3951 | 4012 | 3902 | 3972 | 4002 | 3844 | 3976 | 3654 | 3387 | 3082 | 2790 | 2496 | 2335 | 2032 | 1926 | 1758 | 1560 | 1522 | 1357 | 1320 | 1126 | 1055 | 988 | 865 |
| ta | ta | 133067 | 59.8 | 49 | 1.4e-01 | 2.2 | -0.35 | 5 | 1025 | 1081 | 1152 | 1224 | 1396 | 1463 | 1552 | 1699 | 1869 | 1996 | 2185 | 2409 | 2596 | 2876 | 3028 | 3368 | 3812 | 3994 | 4132 | 3927 | 4080 | 3946 | 3891 | 4045 | 4093 | 4076 | 3940 | 4024 | 3910 | 3927 | 3961 | 3867 | 3941 | 3805 | 3408 | 3093 | 2846 | 2593 | 2321 | 2173 | 1989 | 1814 | 1682 | 1554 | 1446 | 1281 | 1283 | 1227 | 1065 | 1002 |
| at | at | 173691 | 59.2 | 49 | 1.5e-01 | 2.4 | -0.39 | 6 | 1325 | 1426 | 1504 | 1681 | 1793 | 1910 | 2014 | 2181 | 2456 | 2554 | 2755 | 3175 | 3325 | 3678 | 3996 | 4440 | 4974 | 5192 | 5173 | 5248 | 5338 | 5237 | 5115 | 5315 | 5369 | 5243 | 5242 | 5171 | 5194 | 5163 | 5122 | 5202 | 5175 | 5021 | 4525 | 3960 | 3727 | 3402 | 2973 | 2834 | 2560 | 2386 | 2210 | 1954 | 1848 | 1691 | 1707 | 1586 | 1366 | 1255 |
| tc | tc | 150157 | 57.9 | 49 | 1.8e-01 | 2.9 | -0.46 | 7 | 1194 | 1199 | 1307 | 1441 | 1532 | 1680 | 1738 | 1880 | 2111 | 2213 | 2381 | 2703 | 2878 | 3124 | 3431 | 3966 | 4186 | 4671 | 4560 | 4574 | 4596 | 4588 | 4589 | 4526 | 4491 | 4533 | 4652 | 4584 | 4714 | 4567 | 4494 | 4602 | 4537 | 4274 | 3750 | 3356 | 3013 | 2789 | 2522 | 2387 | 2193 | 2017 | 1858 | 1746 | 1610 | 1428 | 1419 | 1289 | 1157 | 1107 |
| ct | ct | 145631 | 53.9 | 49 | 2.9e-01 | 4.7 | -0.67 | 8 | 1171 | 1195 | 1280 | 1362 | 1468 | 1581 | 1720 | 1825 | 2021 | 2126 | 2355 | 2597 | 2851 | 3035 | 3366 | 3843 | 4167 | 4495 | 4543 | 4400 | 4497 | 4406 | 4395 | 4436 | 4274 | 4408 | 4434 | 4423 | 4519 | 4400 | 4347 | 4366 | 4400 | 4189 | 3560 | 3388 | 2974 | 2786 | 2507 | 2235 | 2102 | 1914 | 1729 | 1707 | 1624 | 1392 | 1333 | 1241 | 1188 | 1056 |
| ac | ac | 143481 | 52.3 | 49 | 3.5e-01 | 5.6 | -0.75 | 9 | 1091 | 1198 | 1242 | 1316 | 1443 | 1509 | 1737 | 1852 | 1920 | 2092 | 2348 | 2454 | 2782 | 2982 | 3417 | 3781 | 4195 | 4242 | 4413 | 4365 | 4365 | 4294 | 4465 | 4271 | 4371 | 4380 | 4254 | 4281 | 4221 | 4359 | 4409 | 4423 | 4269 | 4013 | 3602 | 3304 | 2966 | 2723 | 2506 | 2216 | 2138 | 1953 | 1776 | 1746 | 1602 | 1381 | 1325 | 1298 | 1162 | 1029 |
| cg | cg | 141987 | 49.8 | 49 | 4.4e-01 | 7 | -0.85 | 10 | 1035 | 1138 | 1256 | 1308 | 1413 | 1542 | 1634 | 1832 | 1957 | 2125 | 2242 | 2478 | 2729 | 3084 | 3259 | 3718 | 3950 | 4381 | 4213 | 4474 | 4300 | 4413 | 4375 | 4290 | 4364 | 4327 | 4320 | 4401 | 4245 | 4290 | 4388 | 4337 | 4329 | 3896 | 3587 | 3237 | 2947 | 2707 | 2431 | 2233 | 2070 | 1934 | 1769 | 1612 | 1496 | 1390 | 1301 | 1171 | 1087 | 972 |
| gc | gc | 174957 | 46.7 | 49 | 5.7e-01 | 9.1 | -0.96 | 11 | 1297 | 1421 | 1574 | 1637 | 1732 | 1844 | 2081 | 2212 | 2387 | 2634 | 2799 | 3037 | 3351 | 3693 | 4053 | 4481 | 4971 | 5387 | 5280 | 5400 | 5244 | 5371 | 5352 | 5414 | 5296 | 5322 | 5283 | 5456 | 5340 | 5375 | 5322 | 5238 | 5336 | 4831 | 4356 | 4010 | 3708 | 3431 | 3054 | 2689 | 2571 | 2307 | 2175 | 1949 | 1838 | 1776 | 1557 | 1465 | 1402 | 1218 |
| ga | ga | 148634 | 46.1 | 49 | 5.9e-01 | 9.5 | -0.98 | 12 | 1122 | 1272 | 1294 | 1363 | 1449 | 1607 | 1698 | 1880 | 1962 | 2092 | 2365 | 2669 | 2898 | 3186 | 3520 | 3802 | 4236 | 4443 | 4405 | 4560 | 4540 | 4492 | 4536 | 4504 | 4483 | 4608 | 4472 | 4450 | 4442 | 4540 | 4412 | 4568 | 4603 | 4234 | 3868 | 3478 | 3153 | 2835 | 2571 | 2454 | 2134 | 1981 | 1806 | 1676 | 1605 | 1496 | 1383 | 1279 | 1193 | 1015 |
| ag | ag | 146035 | 39.6 | 49 | 8.3e-01 | 13 | -1.12 | 13 | 1111 | 1233 | 1259 | 1369 | 1432 | 1555 | 1722 | 1798 | 1951 | 2136 | 2359 | 2629 | 2822 | 3125 | 3389 | 3703 | 4201 | 4355 | 4434 | 4334 | 4290 | 4383 | 4458 | 4448 | 4352 | 4557 | 4430 | 4495 | 4447 | 4516 | 4367 | 4384 | 4462 | 4062 | 3739 | 3410 | 3081 | 2793 | 2602 | 2378 | 2148 | 1921 | 1808 | 1675 | 1541 | 1528 | 1321 | 1242 | 1216 | 1064 |
| gt | gt | 144318 | 36.2 | 49 | 9.1e-01 | 15 | -1.16 | 14 | 1066 | 1167 | 1286 | 1332 | 1518 | 1584 | 1660 | 1809 | 1933 | 2203 | 2387 | 2596 | 2787 | 3132 | 3329 | 3649 | 4031 | 4370 | 4360 | 4426 | 4266 | 4341 | 4353 | 4383 | 4307 | 4357 | 4423 | 4423 | 4389 | 4310 | 4374 | 4292 | 4344 | 4039 | 3755 | 3354 | 2990 | 2733 | 2508 | 2378 | 2153 | 1931 | 1763 | 1704 | 1501 | 1451 | 1352 | 1263 | 1164 | 1092 |
| ca | ca | 181088 | 34.2 | 49 | 9.5e-01 | 15 | -1.18 | 15 | 1400 | 1468 | 1572 | 1737 | 1821 | 1907 | 2195 | 2282 | 2471 | 2692 | 2893 | 3165 | 3411 | 3725 | 4250 | 4718 | 5205 | 5435 | 5496 | 5471 | 5335 | 5533 | 5590 | 5512 | 5496 | 5475 | 5496 | 5506 | 5470 | 5614 | 5507 | 5565 | 5384 | 5030 | 4568 | 4069 | 3778 | 3452 | 3170 | 2839 | 2685 | 2447 | 2303 | 2121 | 1916 | 1847 | 1688 | 1611 | 1473 | 1294 |
| tg | tg | 180048 | 31.6 | 49 | 9.7e-01 | 16 | -1.19 | 16 | 1339 | 1500 | 1613 | 1697 | 1838 | 1918 | 2091 | 2223 | 2400 | 2690 | 2928 | 3189 | 3521 | 3837 | 4226 | 4540 | 5122 | 5399 | 5475 | 5501 | 5429 | 5486 | 5434 | 5522 | 5385 | 5381 | 5487 | 5414 | 5461 | 5392 | 5342 | 5409 | 5504 | 5119 | 4686 | 4200 | 3801 | 3538 | 3147 | 2889 | 2616 | 2367 | 2209 | 2021 | 1915 | 1833 | 1670 | 1570 | 1500 | 1274 |
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