position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/02/26/peak-motifs.2026-02-26.200629_2026-02-26.200629_uoww0v/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title SMAD2_hela_hg38_SEACR_relaxed -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/02/26/peak-motifs.2026-02-26.200629_2026-02-26.200629_uoww0v/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/02/26/peak-motifs.2026-02-26.200629_2026-02-26.200629_uoww0v/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/02/26/peak-motifs.2026-02-26.200629_2026-02-26.200629_uoww0v/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 226 Sum of sequence lengths 226000 Min sequence length 0 Max sequence length 1000 Average sequence length 1000 Possible positions 225774 Oligonucleotide statistics: Total occurrences 196214 Position interval parameters: Position interval 20 Number of windows 50 Total positions 225774 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 226 4520 2 -24 [-479,-460] -469.5 226 4520 3 -23 [-459,-440] -449.5 226 4520 4 -22 [-439,-420] -429.5 226 4520 5 -21 [-419,-400] -409.5 226 4520 6 -20 [-399,-380] -389.5 226 4520 7 -19 [-379,-360] -369.5 226 4520 8 -18 [-359,-340] -349.5 226 4520 9 -17 [-339,-320] -329.5 226 4520 10 -16 [-319,-300] -309.5 226 4520 11 -15 [-299,-280] -289.5 226 4520 12 -14 [-279,-260] -269.5 226 4520 13 -13 [-259,-240] -249.5 226 4520 14 -12 [-239,-220] -229.5 226 4520 15 -11 [-219,-200] -209.5 226 4520 16 -10 [-199,-180] -189.5 226 4520 17 -9 [-179,-160] -169.5 226 4520 18 -8 [-159,-140] -149.5 226 4520 19 -7 [-139,-120] -129.5 226 4520 20 -6 [-119,-100] -109.5 226 4520 21 -5 [-99,-80] -89.5 226 4520 22 -4 [-79,-60] -69.5 226 4520 23 -3 [-59,-40] -49.5 226 4520 24 -2 [-39,-20] -29.5 226 4520 25 -1 [-19,0] -9.5 226 4520 26 0 [1,20] 10.5 226 4520 27 1 [21,40] 30.5 226 4520 28 2 [41,60] 50.5 226 4520 29 3 [61,80] 70.5 226 4520 30 4 [81,100] 90.5 226 4520 31 5 [101,120] 110.5 226 4520 32 6 [121,140] 130.5 226 4520 33 7 [141,160] 150.5 226 4520 34 8 [161,180] 170.5 226 4520 35 9 [181,200] 190.5 226 4520 36 10 [201,220] 210.5 226 4520 37 11 [221,240] 230.5 226 4520 38 12 [241,260] 250.5 226 4520 39 13 [261,280] 270.5 226 4520 40 14 [281,300] 290.5 226 4520 41 15 [301,320] 310.5 226 4520 42 16 [321,340] 330.5 226 4520 43 17 [341,360] 350.5 226 4520 44 18 [361,380] 370.5 226 4520 45 19 [381,400] 390.5 226 4520 46 20 [401,420] 410.5 226 4520 47 21 [421,440] 430.5 226 4520 48 22 [441,460] 450.5 226 4520 49 23 [461,480] 470.5 226 4520 50 24 [481,500] 490.5 226 4294 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| cc | cc | 10092 | 97.4 | 49 | 4.7e-05 | 0.00076 | 3.12 | 1 | 242 | 204 | 177 | 181 | 202 | 207 | 211 | 226 | 185 | 179 | 209 | 217 | 208 | 164 | 224 | 210 | 205 | 178 | 204 | 215 | 172 | 221 | 228 | 189 | 213 | 215 | 210 | 211 | 229 | 188 | 194 | 207 | 202 | 235 | 229 | 196 | 228 | 218 | 189 | 181 | 209 | 188 | 184 | 195 | 218 | 142 | 175 | 185 | 199 | 194 |
| tt | tt | 19585 | 94.9 | 49 | 9.2e-05 | 0.0015 | 2.83 | 2 | 389 | 357 | 381 | 360 | 357 | 394 | 393 | 388 | 375 | 395 | 391 | 348 | 396 | 375 | 374 | 373 | 423 | 373 | 413 | 376 | 404 | 388 | 377 | 375 | 365 | 401 | 413 | 401 | 365 | 439 | 416 | 441 | 440 | 380 | 339 | 388 | 337 | 363 | 406 | 436 | 403 | 442 | 424 | 396 | 440 | 441 | 411 | 384 | 383 | 356 |
| gg | gg | 9746 | 80.5 | 49 | 3.0e-03 | 0.049 | 1.31 | 3 | 205 | 179 | 193 | 246 | 206 | 208 | 202 | 177 | 192 | 203 | 178 | 186 | 191 | 186 | 194 | 206 | 192 | 215 | 187 | 160 | 225 | 183 | 202 | 198 | 192 | 209 | 181 | 177 | 211 | 173 | 230 | 210 | 181 | 206 | 200 | 178 | 193 | 171 | 179 | 183 | 219 | 193 | 203 | 222 | 182 | 154 | 212 | 195 | 177 | 201 |
| ct | ct | 14007 | 77.7 | 49 | 5.6e-03 | 0.09 | 1.05 | 4 | 287 | 283 | 253 | 244 | 279 | 247 | 290 | 296 | 291 | 264 | 262 | 265 | 288 | 288 | 296 | 295 | 270 | 275 | 293 | 300 | 283 | 295 | 311 | 291 | 269 | 286 | 277 | 309 | 281 | 315 | 315 | 319 | 309 | 287 | 268 | 240 | 256 | 281 | 298 | 255 | 261 | 277 | 300 | 267 | 253 | 262 | 256 | 258 | 254 | 308 |
| aa | aa | 19526 | 70.8 | 49 | 2.3e-02 | 0.36 | 0.44 | 5 | 377 | 435 | 388 | 387 | 364 | 412 | 388 | 361 | 416 | 405 | 387 | 411 | 399 | 409 | 368 | 427 | 359 | 378 | 343 | 396 | 440 | 392 | 431 | 385 | 371 | 384 | 365 | 447 | 408 | 391 | 353 | 375 | 376 | 396 | 413 | 361 | 400 | 403 | 375 | 406 | 409 | 391 | 366 | 413 | 358 | 377 | 367 | 376 | 403 | 384 |
| ac | ac | 9760 | 64.5 | 49 | 6.8e-02 | 1.1 | -0.04 | 6 | 185 | 174 | 208 | 190 | 231 | 168 | 163 | 211 | 188 | 183 | 207 | 209 | 209 | 189 | 201 | 201 | 186 | 207 | 216 | 201 | 177 | 202 | 212 | 192 | 215 | 182 | 197 | 188 | 184 | 213 | 191 | 199 | 209 | 206 | 221 | 208 | 177 | 198 | 188 | 176 | 200 | 208 | 176 | 201 | 213 | 194 | 153 | 189 | 171 | 193 |
| ta | ta | 12890 | 59.8 | 49 | 1.4e-01 | 2.2 | -0.35 | 7 | 248 | 239 | 263 | 228 | 270 | 277 | 246 | 248 | 238 | 249 | 250 | 234 | 260 | 254 | 265 | 238 | 238 | 252 | 285 | 265 | 263 | 301 | 229 | 254 | 234 | 262 | 260 | 266 | 281 | 268 | 267 | 252 | 287 | 251 | 243 | 252 | 239 | 265 | 258 | 277 | 253 | 273 | 248 | 233 | 273 | 304 | 288 | 252 | 253 | 257 |
| ca | ca | 14183 | 56.7 | 49 | 2.1e-01 | 3.4 | -0.53 | 8 | 286 | 282 | 276 | 292 | 305 | 279 | 239 | 287 | 274 | 273 | 313 | 315 | 315 | 279 | 301 | 276 | 283 | 286 | 287 | 308 | 262 | 274 | 290 | 272 | 314 | 254 | 295 | 253 | 279 | 295 | 259 | 274 | 291 | 288 | 312 | 318 | 292 | 286 | 265 | 268 | 282 | 285 | 254 | 298 | 298 | 268 | 258 | 285 | 292 | 266 |
| gt | gt | 9866 | 51.1 | 49 | 3.9e-01 | 6.2 | -0.79 | 9 | 186 | 182 | 194 | 228 | 200 | 170 | 199 | 174 | 174 | 198 | 190 | 202 | 182 | 212 | 203 | 188 | 199 | 219 | 185 | 200 | 207 | 195 | 175 | 226 | 207 | 196 | 186 | 185 | 203 | 202 | 207 | 217 | 194 | 213 | 195 | 193 | 194 | 191 | 221 | 198 | 202 | 174 | 216 | 178 | 206 | 212 | 221 | 201 | 195 | 171 |
| gc | gc | 8149 | 49.0 | 49 | 4.7e-01 | 7.6 | -0.88 | 10 | 173 | 172 | 145 | 171 | 155 | 160 | 148 | 177 | 166 | 154 | 159 | 174 | 165 | 163 | 191 | 182 | 146 | 169 | 165 | 189 | 159 | 161 | 158 | 167 | 180 | 151 | 155 | 158 | 187 | 147 | 176 | 177 | 176 | 160 | 166 | 165 | 150 | 163 | 143 | 150 | 143 | 160 | 157 | 147 | 154 | 143 | 157 | 168 | 187 | 160 |
| tc | tc | 11935 | 47.4 | 49 | 5.4e-01 | 8.6 | -0.93 | 11 | 262 | 246 | 226 | 223 | 227 | 222 | 260 | 230 | 251 | 230 | 240 | 248 | 250 | 244 | 247 | 223 | 252 | 217 | 247 | 236 | 242 | 231 | 255 | 248 | 229 | 247 | 244 | 248 | 237 | 265 | 254 | 244 | 248 | 239 | 222 | 220 | 256 | 249 | 266 | 225 | 233 | 216 | 239 | 255 | 216 | 219 | 230 | 204 | 219 | 254 |
| cg | cg | 1656 | 44.9 | 49 | 6.4e-01 | 10 | -1.01 | 12 | 40 | 39 | 39 | 41 | 40 | 27 | 34 | 40 | 29 | 29 | 41 | 42 | 23 | 32 | 37 | 37 | 30 | 34 | 34 | 32 | 35 | 24 | 30 | 39 | 39 | 37 | 36 | 34 | 34 | 30 | 34 | 39 | 26 | 31 | 28 | 33 | 34 | 46 | 36 | 31 | 29 | 28 | 30 | 24 | 33 | 28 | 21 | 30 | 27 | 30 |
| tg | tg | 14204 | 42.9 | 49 | 7.2e-01 | 11 | -1.06 | 13 | 281 | 263 | 287 | 301 | 272 | 253 | 294 | 259 | 280 | 295 | 270 | 280 | 273 | 311 | 289 | 296 | 293 | 319 | 280 | 308 | 305 | 277 | 268 | 308 | 288 | 290 | 260 | 279 | 276 | 279 | 306 | 306 | 292 | 303 | 283 | 276 | 273 | 261 | 291 | 284 | 268 | 274 | 305 | 264 | 259 | 283 | 284 | 301 | 289 | 268 |
| ag | ag | 13807 | 40.3 | 49 | 8.1e-01 | 13 | -1.11 | 14 | 299 | 288 | 289 | 288 | 295 | 290 | 259 | 271 | 269 | 276 | 269 | 296 | 279 | 276 | 272 | 282 | 277 | 272 | 263 | 291 | 280 | 265 | 262 | 276 | 291 | 251 | 275 | 244 | 320 | 276 | 290 | 276 | 277 | 270 | 292 | 259 | 260 | 267 | 276 | 247 | 275 | 249 | 256 | 271 | 275 | 279 | 286 | 292 | 302 | 267 |
| ga | ga | 11653 | 35.7 | 49 | 9.2e-01 | 15 | -1.17 | 15 | 265 | 238 | 261 | 248 | 251 | 234 | 229 | 234 | 247 | 238 | 229 | 254 | 218 | 245 | 206 | 238 | 252 | 237 | 237 | 234 | 241 | 227 | 218 | 231 | 231 | 234 | 228 | 225 | 224 | 236 | 252 | 231 | 224 | 229 | 233 | 220 | 215 | 231 | 225 | 227 | 233 | 211 | 212 | 230 | 216 | 225 | 220 | 245 | 241 | 243 |
| at | at | 15155 | 32.9 | 49 | 9.6e-01 | 15 | -1.19 | 16 | 314 | 298 | 299 | 294 | 291 | 331 | 312 | 275 | 306 | 293 | 312 | 303 | 312 | 303 | 294 | 296 | 306 | 297 | 333 | 308 | 311 | 317 | 280 | 285 | 282 | 305 | 322 | 295 | 294 | 301 | 300 | 283 | 307 | 305 | 279 | 311 | 319 | 307 | 299 | 325 | 296 | 315 | 280 | 303 | 296 | 340 | 313 | 293 | 319 | 296 |
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