position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/02/26/peak-motifs.2026-02-26.211242_2026-02-26.211242_nt22t5/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title SMAD2_helasherg_SEACR_stringent -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/02/26/peak-motifs.2026-02-26.211242_2026-02-26.211242_nt22t5/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/02/26/peak-motifs.2026-02-26.211242_2026-02-26.211242_nt22t5/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/02/26/peak-motifs.2026-02-26.211242_2026-02-26.211242_nt22t5/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 342 Sum of sequence lengths 341706 Min sequence length 0 Max sequence length 1000 Average sequence length 999 Possible positions 341364 Oligonucleotide statistics: Total occurrences 284746 Position interval parameters: Position interval 20 Number of windows 50 Total positions 341364 Degrees of freedom 49 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -25 [-499,-480] -489.5 341 6820 2 -24 [-479,-460] -469.5 341 6820 3 -23 [-459,-440] -449.5 341 6820 4 -22 [-439,-420] -429.5 341 6820 5 -21 [-419,-400] -409.5 341 6820 6 -20 [-399,-380] -389.5 341 6820 7 -19 [-379,-360] -369.5 341 6820 8 -18 [-359,-340] -349.5 342 6834 9 -17 [-339,-320] -329.5 342 6840 10 -16 [-319,-300] -309.5 342 6840 11 -15 [-299,-280] -289.5 342 6840 12 -14 [-279,-260] -269.5 342 6840 13 -13 [-259,-240] -249.5 342 6840 14 -12 [-239,-220] -229.5 342 6840 15 -11 [-219,-200] -209.5 342 6840 16 -10 [-199,-180] -189.5 342 6840 17 -9 [-179,-160] -169.5 342 6840 18 -8 [-159,-140] -149.5 342 6840 19 -7 [-139,-120] -129.5 342 6840 20 -6 [-119,-100] -109.5 342 6840 21 -5 [-99,-80] -89.5 342 6840 22 -4 [-79,-60] -69.5 342 6840 23 -3 [-59,-40] -49.5 342 6840 24 -2 [-39,-20] -29.5 342 6840 25 -1 [-19,0] -9.5 342 6840 26 0 [1,20] 10.5 342 6840 27 1 [21,40] 30.5 342 6840 28 2 [41,60] 50.5 342 6840 29 3 [61,80] 70.5 342 6840 30 4 [81,100] 90.5 342 6840 31 5 [101,120] 110.5 342 6840 32 6 [121,140] 130.5 342 6840 33 7 [141,160] 150.5 342 6840 34 8 [161,180] 170.5 342 6840 35 9 [181,200] 190.5 342 6840 36 10 [201,220] 210.5 342 6840 37 11 [221,240] 230.5 342 6840 38 12 [241,260] 250.5 342 6840 39 13 [261,280] 270.5 342 6840 40 14 [281,300] 290.5 342 6840 41 15 [301,320] 310.5 342 6840 42 16 [321,340] 330.5 342 6840 43 17 [341,360] 350.5 342 6831 44 18 [361,380] 370.5 341 6820 45 19 [381,400] 390.5 341 6820 46 20 [401,420] 410.5 341 6820 47 21 [421,440] 430.5 341 6820 48 22 [441,460] 450.5 341 6820 49 23 [461,480] 470.5 341 6820 50 24 [481,500] 490.5 341 6479 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -489.5 occurrences in window 1 [-499,-480] 11 -469.5 occurrences in window 2 [-479,-460] 12 -449.5 occurrences in window 3 [-459,-440] 13 -429.5 occurrences in window 4 [-439,-420] 14 -409.5 occurrences in window 5 [-419,-400] 15 -389.5 occurrences in window 6 [-399,-380] 16 -369.5 occurrences in window 7 [-379,-360] 17 -349.5 occurrences in window 8 [-359,-340] 18 -329.5 occurrences in window 9 [-339,-320] 19 -309.5 occurrences in window 10 [-319,-300] 20 -289.5 occurrences in window 11 [-299,-280] 21 -269.5 occurrences in window 12 [-279,-260] 22 -249.5 occurrences in window 13 [-259,-240] 23 -229.5 occurrences in window 14 [-239,-220] 24 -209.5 occurrences in window 15 [-219,-200] 25 -189.5 occurrences in window 16 [-199,-180] 26 -169.5 occurrences in window 17 [-179,-160] 27 -149.5 occurrences in window 18 [-159,-140] 28 -129.5 occurrences in window 19 [-139,-120] 29 -109.5 occurrences in window 20 [-119,-100] 30 -89.5 occurrences in window 21 [-99,-80] 31 -69.5 occurrences in window 22 [-79,-60] 32 -49.5 occurrences in window 23 [-59,-40] 33 -29.5 occurrences in window 24 [-39,-20] 34 -9.5 occurrences in window 25 [-19,0] 35 10.5 occurrences in window 26 [1,20] 36 30.5 occurrences in window 27 [21,40] 37 50.5 occurrences in window 28 [41,60] 38 70.5 occurrences in window 29 [61,80] 39 90.5 occurrences in window 30 [81,100] 40 110.5 occurrences in window 31 [101,120] 41 130.5 occurrences in window 32 [121,140] 42 150.5 occurrences in window 33 [141,160] 43 170.5 occurrences in window 34 [161,180] 44 190.5 occurrences in window 35 [181,200] 45 210.5 occurrences in window 36 [201,220] 46 230.5 occurrences in window 37 [221,240] 47 250.5 occurrences in window 38 [241,260] 48 270.5 occurrences in window 39 [261,280] 49 290.5 occurrences in window 40 [281,300] 50 310.5 occurrences in window 41 [301,320] 51 330.5 occurrences in window 42 [321,340] 52 350.5 occurrences in window 43 [341,360] 53 370.5 occurrences in window 44 [361,380] 54 390.5 occurrences in window 45 [381,400] 55 410.5 occurrences in window 46 [401,420] 56 430.5 occurrences in window 47 [421,440] 57 450.5 occurrences in window 48 [441,460] 58 470.5 occurrences in window 49 [461,480] 59 490.5 occurrences in window 50 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| tt | tt | 28002 | 98.1 | 49 | 3.9e-05 | 0.00063 | 3.20 | 1 | 563 | 577 | 570 | 538 | 526 | 570 | 560 | 596 | 587 | 591 | 610 | 578 | 633 | 629 | 567 | 562 | 606 | 573 | 572 | 519 | 497 | 530 | 558 | 580 | 595 | 577 | 644 | 527 | 533 | 556 | 562 | 528 | 510 | 519 | 510 | 590 | 554 | 537 | 526 | 514 | 567 | 577 | 582 | 559 | 538 | 520 | 525 | 542 | 563 | 555 |
| gg | gg | 14221 | 88.3 | 49 | 4.9e-04 | 0.0078 | 2.11 | 2 | 320 | 323 | 321 | 321 | 292 | 305 | 304 | 305 | 307 | 294 | 287 | 302 | 304 | 266 | 222 | 272 | 242 | 267 | 260 | 267 | 276 | 278 | 271 | 276 | 257 | 294 | 277 | 298 | 289 | 289 | 251 | 297 | 282 | 273 | 298 | 279 | 272 | 294 | 281 | 329 | 255 | 253 | 272 | 281 | 272 | 291 | 270 | 322 | 297 | 266 |
| ga | ga | 17149 | 81.3 | 49 | 2.5e-03 | 0.041 | 1.39 | 3 | 371 | 398 | 368 | 361 | 375 | 358 | 395 | 392 | 356 | 331 | 364 | 377 | 377 | 344 | 354 | 344 | 323 | 334 | 333 | 335 | 345 | 332 | 313 | 366 | 341 | 321 | 308 | 321 | 337 | 304 | 344 | 321 | 330 | 336 | 317 | 315 | 328 | 376 | 328 | 322 | 307 | 339 | 335 | 339 | 345 | 323 | 319 | 351 | 346 | 350 |
| ag | ag | 20016 | 79.0 | 49 | 4.2e-03 | 0.067 | 1.17 | 4 | 463 | 441 | 434 | 423 | 410 | 407 | 441 | 467 | 401 | 391 | 415 | 435 | 428 | 416 | 417 | 381 | 407 | 382 | 385 | 372 | 426 | 391 | 377 | 390 | 384 | 372 | 366 | 406 | 360 | 362 | 380 | 395 | 378 | 386 | 360 | 384 | 392 | 430 | 406 | 392 | 384 | 380 | 378 | 409 | 395 | 397 | 396 | 404 | 423 | 397 |
| aa | aa | 28901 | 69.4 | 49 | 2.9e-02 | 0.47 | 0.33 | 5 | 611 | 619 | 603 | 556 | 548 | 581 | 587 | 576 | 590 | 579 | 592 | 533 | 618 | 587 | 600 | 580 | 603 | 568 | 569 | 560 | 565 | 557 | 521 | 574 | 624 | 578 | 540 | 576 | 602 | 581 | 573 | 586 | 607 | 552 | 588 | 547 | 540 | 552 | 560 | 551 | 623 | 603 | 558 | 612 | 538 | 576 | 539 | 567 | 644 | 607 |
| cc | cc | 13800 | 66.1 | 49 | 5.2e-02 | 0.84 | 0.08 | 6 | 276 | 274 | 296 | 258 | 293 | 293 | 262 | 253 | 256 | 287 | 277 | 272 | 299 | 273 | 275 | 312 | 290 | 278 | 255 | 291 | 257 | 290 | 294 | 274 | 251 | 272 | 273 | 271 | 282 | 268 | 279 | 275 | 303 | 221 | 275 | 284 | 302 | 258 | 256 | 238 | 267 | 276 | 249 | 295 | 283 | 305 | 291 | 263 | 272 | 306 |
| ac | ac | 14472 | 57.3 | 49 | 1.9e-01 | 3.1 | -0.49 | 7 | 317 | 286 | 295 | 281 | 325 | 298 | 284 | 289 | 285 | 306 | 286 | 271 | 282 | 258 | 288 | 322 | 315 | 281 | 292 | 302 | 280 | 287 | 318 | 260 | 283 | 259 | 288 | 290 | 280 | 253 | 291 | 295 | 300 | 304 | 309 | 258 | 279 | 281 | 261 | 310 | 315 | 258 | 317 | 287 | 292 | 287 | 306 | 291 | 276 | 294 |
| ta | ta | 19591 | 55.6 | 49 | 2.4e-01 | 3.9 | -0.59 | 8 | 422 | 357 | 378 | 392 | 400 | 366 | 387 | 379 | 375 | 390 | 426 | 410 | 345 | 405 | 398 | 411 | 397 | 397 | 378 | 396 | 385 | 385 | 406 | 388 | 423 | 389 | 411 | 370 | 368 | 362 | 345 | 401 | 367 | 430 | 387 | 389 | 398 | 379 | 403 | 391 | 413 | 373 | 437 | 387 | 421 | 411 | 368 | 395 | 406 | 394 |
| gc | gc | 11575 | 54.6 | 49 | 2.7e-01 | 4.3 | -0.64 | 9 | 257 | 231 | 227 | 234 | 245 | 251 | 253 | 236 | 257 | 238 | 214 | 223 | 229 | 225 | 233 | 213 | 213 | 260 | 223 | 235 | 204 | 230 | 221 | 207 | 193 | 216 | 223 | 229 | 226 | 234 | 210 | 255 | 233 | 241 | 256 | 257 | 244 | 232 | 214 | 241 | 227 | 219 | 213 | 229 | 217 | 246 | 259 | 239 | 234 | 229 |
| tc | tc | 16560 | 51.8 | 49 | 3.6e-01 | 5.8 | -0.76 | 10 | 327 | 330 | 338 | 367 | 336 | 365 | 329 | 336 | 313 | 319 | 322 | 337 | 326 | 339 | 384 | 352 | 337 | 299 | 331 | 315 | 318 | 319 | 346 | 343 | 310 | 350 | 320 | 314 | 312 | 323 | 343 | 334 | 327 | 301 | 309 | 316 | 355 | 334 | 346 | 308 | 331 | 346 | 306 | 327 | 365 | 334 | 346 | 302 | 334 | 339 |
| cg | cg | 2374 | 50.2 | 49 | 4.2e-01 | 6.8 | -0.83 | 11 | 61 | 55 | 50 | 48 | 51 | 60 | 48 | 42 | 47 | 47 | 50 | 45 | 53 | 46 | 56 | 49 | 36 | 52 | 41 | 51 | 37 | 51 | 50 | 48 | 32 | 48 | 53 | 26 | 52 | 42 | 47 | 56 | 53 | 57 | 55 | 53 | 54 | 42 | 44 | 47 | 46 | 47 | 46 | 44 | 48 | 45 | 43 | 43 | 31 | 46 |
| ct | ct | 19842 | 49.9 | 49 | 4.4e-01 | 7 | -0.84 | 12 | 415 | 402 | 406 | 430 | 438 | 419 | 421 | 400 | 392 | 413 | 382 | 398 | 367 | 379 | 422 | 428 | 396 | 402 | 395 | 380 | 372 | 389 | 413 | 384 | 368 | 392 | 384 | 375 | 355 | 363 | 378 | 400 | 405 | 397 | 401 | 401 | 399 | 392 | 405 | 390 | 402 | 398 | 350 | 413 | 443 | 388 | 417 | 395 | 410 | 378 |
| tg | tg | 20772 | 48.0 | 49 | 5.1e-01 | 8.2 | -0.91 | 13 | 409 | 440 | 407 | 428 | 437 | 430 | 442 | 414 | 459 | 453 | 396 | 424 | 389 | 414 | 398 | 414 | 400 | 427 | 432 | 391 | 419 | 397 | 371 | 388 | 407 | 406 | 404 | 413 | 390 | 417 | 429 | 410 | 409 | 440 | 418 | 429 | 406 | 404 | 417 | 430 | 369 | 434 | 406 | 419 | 410 | 405 | 415 | 467 | 427 | 412 |
| gt | gt | 14422 | 48.0 | 49 | 5.1e-01 | 8.2 | -0.91 | 14 | 294 | 310 | 299 | 307 | 276 | 280 | 290 | 292 | 308 | 318 | 276 | 304 | 267 | 305 | 286 | 291 | 285 | 295 | 292 | 253 | 308 | 293 | 266 | 248 | 286 | 292 | 284 | 279 | 252 | 298 | 283 | 294 | 275 | 301 | 270 | 285 | 264 | 294 | 306 | 310 | 275 | 295 | 287 | 293 | 285 | 287 | 269 | 329 | 302 | 284 |
| at | at | 22656 | 42.1 | 49 | 7.5e-01 | 12 | -1.08 | 15 | 444 | 425 | 415 | 460 | 454 | 462 | 457 | 438 | 449 | 433 | 493 | 445 | 435 | 478 | 463 | 457 | 451 | 441 | 457 | 461 | 425 | 438 | 448 | 475 | 486 | 474 | 453 | 434 | 478 | 445 | 446 | 438 | 443 | 461 | 442 | 454 | 475 | 448 | 452 | 421 | 446 | 459 | 489 | 423 | 470 | 480 | 446 | 444 | 458 | 487 |
| ca | ca | 20393 | 39.5 | 49 | 8.3e-01 | 13 | -1.12 | 16 | 415 | 394 | 406 | 410 | 412 | 433 | 408 | 411 | 408 | 408 | 406 | 367 | 418 | 402 | 409 | 408 | 437 | 397 | 415 | 416 | 392 | 403 | 413 | 385 | 378 | 403 | 390 | 421 | 423 | 409 | 416 | 411 | 406 | 390 | 417 | 387 | 418 | 401 | 398 | 398 | 439 | 384 | 433 | 383 | 390 | 427 | 462 | 390 | 421 | 425 |
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