RSAT - peak-motifs result

Analysis: SMAD2_helasherg_SEACR_stringent_peak_motifs (19/03/2026 14:47)

peak-motifs  -v 1 -title SMAD2_helasherg_SEACR_stringent_peak_motifs -i $RSAT/public_html/tmp/www-data/2026/03/19/peak-motifs.2026-03-19.143247_2026-03-19.143247_dTipSp/peak-motifspeak_seq -max_seq_len 1000 -markov auto -disco oligos,dyads,positions,local_words -nmotifs 10 -minol 6 -maxol 8 -no_merge_lengths -2str -origin center -motif_db JASPAR2024_CORE_vertebrates_non-redundant tf $RSAT/public_html/motif_databases/JASPAR/Jaspar_2024/JASPAR2024_CORE_vertebrates_non-redundant_pfms_transfac.txt -scan_markov 1 -task purge,seqlen,composition,disco,merge_motifs,split_motifs,motifs_vs_motifs,timelog,archive,synthesis,small_summary,motifs_vs_db,scan -prefix peak-motifs -noov -img_format png -outdir $RSAT/public_html/tmp/www-data/2026/03/19/peak-motifs.2026-03-19.143247_2026-03-19.143247_dTipSp

motif discovery
oligos_6nt_mkv3	oligos_6nt_mkv3_m1 assembly: cccagcct (sig=5.57; e-value=2.7e-06) RC: aggctggg oligos_6nt_mkv3_m2 assembly: gcctggc (sig=5.50; e-value=3.2e-06) RC: gccaggc oligos_6nt_mkv3_m3 assembly: aaacaaa (sig=5.23; e-value=5.9e-06) RC: tttgttt oligos_6nt_mkv3_m4 assembly: aaatta (sig=3.86; e-value=0.00014) RC: taattt oligos_6nt_mkv3_m5 assembly: cctccc (sig=3.45; e-value=0.00035) RC: gggagg oligos_6nt_mkv3_m6 assembly: tacata (sig=3.32; e-value=0.00048) RC: tatgta oligos_6nt_mkv3_m7 assembly: aaagaaa (sig=3.21; e-value=0.00062) RC: tttcttt oligos_6nt_mkv3_m8 assembly: aataaa (sig=2.79; e-value=0.0016) RC: tttatt oligos_6nt_mkv3_m9 assembly: gacagagc (sig=2.68; e-value=0.0021) RC: gctctgtc oligos_6nt_mkv3_m10 assembly: gccaagg (sig=2.41; e-value=0.0039) RC: ccttggc	[ discovered words: text ] [ assembly: text sig matrices ] [ matrices: tab format transfac format ]
local_words_6nt	Not a single significant pattern	[local_words_6nt]
positions_6nt	positions_6nt_m1 assembly: agccaa (sig=0.75; e-value=0.18) RC: ttggct positions_6nt_m2 assembly: acttct (sig=0.07; e-value=0.85) RC: agaagt positions_6nt_m3 : (sig=; e-value=1) RC: positions_6nt_m4 : (sig=; e-value=1) RC: positions_6nt_m5 : (sig=; e-value=1) RC: positions_6nt_m6 : (sig=; e-value=1) RC: positions_6nt_m7 : (sig=; e-value=1) RC: positions_6nt_m8 : (sig=; e-value=1) RC: positions_6nt_m9 : (sig=; e-value=1) RC: positions_6nt_m10 : (sig=; e-value=1) RC:	[ discovered words: text ] [ assembly: text sig matrices ] [ matrices: tab format transfac format ]
oligos_7nt_mkv4	oligos_7nt_mkv4_m1 assembly: aaaatta (sig=3.93; e-value=0.00012) RC: taatttt oligos_7nt_mkv4_m2 assembly: aggctgg (sig=2.67; e-value=0.0021) RC: ccagcct oligos_7nt_mkv4_m3 assembly: aaaacaa (sig=1.61; e-value=0.025) RC: ttgtttt oligos_7nt_mkv4_m4 assembly: aaggcaa (sig=1.51; e-value=0.031) RC: ttgcctt oligos_7nt_mkv4_m5 assembly: cagcatc (sig=1.08; e-value=0.083) RC: gatgctg oligos_7nt_mkv4_m6 assembly: gggatta (sig=1.03; e-value=0.093) RC: taatccc oligos_7nt_mkv4_m7 assembly: gcctggcc (sig=1.03; e-value=0.093) RC: ggccaggc oligos_7nt_mkv4_m8 assembly: aatgcaa (sig=0.46; e-value=0.35) RC: ttgcatt oligos_7nt_mkv4_m9 assembly: gaaaagaa (sig=0.21; e-value=0.62) RC: ttcttttc oligos_7nt_mkv4_m10 assembly: aaaggaa (sig=0.07; e-value=0.85) RC: ttccttt	[ discovered words: text ] [ assembly: text sig matrices ] [ matrices: tab format transfac format ]
local_words_7nt	Not a single significant pattern	[local_words_7nt]
positions_7nt	Not a single significant pattern	[positions_7nt]
oligos_8nt_mkv5	oligos_8nt_mkv5_m1 assembly: cctcctcc (sig=3.78; e-value=0.00017) RC: ggaggagg oligos_8nt_mkv5_m2 assembly: acaaaaac (sig=1.77; e-value=0.017) RC: gtttttgt oligos_8nt_mkv5_m3 : (sig=; e-value=1) RC: oligos_8nt_mkv5_m4 : (sig=; e-value=1) RC: oligos_8nt_mkv5_m5 : (sig=; e-value=1) RC: oligos_8nt_mkv5_m6 : (sig=; e-value=1) RC: oligos_8nt_mkv5_m7 : (sig=; e-value=1) RC: oligos_8nt_mkv5_m8 : (sig=; e-value=1) RC: oligos_8nt_mkv5_m9 : (sig=; e-value=1) RC: oligos_8nt_mkv5_m10 : (sig=; e-value=1) RC:	[ discovered words: text ] [ assembly: text sig matrices ] [ matrices: tab format transfac format ]
local_words_8nt	Not a single significant pattern	[local_words_8nt]
positions_8nt	Not a single significant pattern	[positions_8nt]
dyads	dyads_m1 assembly: tatttattttaa (sig=39.36; e-value=4.4e-40) RC: ttaaaataaata dyads_m2 assembly: cctccccccccccagccctc (sig=32.32; e-value=4.8e-33) RC: gagggctggggggggggagg dyads_m3 assembly: ttaaaaataaata (sig=35.65; e-value=2.2e-36) RC: tatttatttttaa dyads_m4 assembly: gcctggc (sig=26.31; e-value=4.9e-27) RC: gccaggc dyads_m5 assembly: cccaggc (sig=20.43; e-value=3.7e-21) RC: gcctggg dyads_m6 assembly: cccacc (sig=19.21; e-value=6.2e-20) RC: ggtggg dyads_m7 assembly: ccccnnctc (sig=18.62; e-value=2.4e-19) RC: gagnngggg dyads_m8 assembly: ttaaaaagaaataat (sig=27.42; e-value=3.8e-28) RC: attatttctttttaa dyads_m9 assembly: gcccccctccccctccctc (sig=18.62; e-value=2.4e-19) RC: gagggagggggaggggggc dyads_m10 assembly: cgcnnngcccg (sig=13.89; e-value=1.3e-14) RC: cgggcnnngcg	[ discovered words: text ] [ assembly: text sig matrices ] [ matrices: tab format transfac format ]

Motif discovery
Motif 1   oligos_6nt_mkv3_m1	k-mer sig=5.57; evalue=2.7e-06 vsCCCAGCCtss SSAGGCTGGGSB	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor vsCCCAGCCtss FEZF2 MA2341.1 D 8 0.6667 0.991 0.661 NA NA Zfp809 MA2125.1 D 9 0.7500 0.881 0.661 NA NA ZNF213 MA2121.1 D 11 0.8462 0.769 0.650 NA NA Total matches= 7 (4 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 33 (9.65%) Max nb of predicted sites per peak: 34  (hg38_ct_MyTrack_6202_15) Mean nb of predicted sites per peak: 17.67	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 2   oligos_6nt_mkv3_m2	k-mer sig=5.50; evalue=3.2e-06 smGCCtGGCys SRGCCAGGCKS	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor smGCCtGGCys ZNF213 MA2121.1 R 10 0.7692 0.761 0.586 NA NA Hand1::Tcf3 MA0092.2 D 9 0.8182 0.706 0.578 NA NA Zfp335 MA2002.2 R 7 0.6364 0.757 0.482 NA NA Total matches= 5 (2 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 31 (9.06%) Max nb of predicted sites per peak: 15  (hg38_ct_MyTrack_6202_15) Mean nb of predicted sites per peak: 9.16	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 3   oligos_6nt_mkv3_m3	k-mer sig=5.23; evalue=5.9e-06 awAAACAAAwa TWTTTGTTTWT	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor awAAACAAAwa FOXD2 MA0847.4 D 11 1.0000 0.790 0.790 NA NA Foxj3 MA0851.2 D 9 0.8182 0.865 0.707 NA NA FOXE1 MA1487.3 D 11 0.9167 0.717 0.657 NA NA Total matches= 41 (38 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 34 (9.94%) Max nb of predicted sites per peak: 8  (hg38_ct_MyTrack_6202_3) Mean nb of predicted sites per peak: 7.24	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 4   oligos_6nt_mkv3_m4	k-mer sig=3.86; evalue=0.00014 aaAAATTArc GYTAATTTTT	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor aaAAATTArc PAX4 MA0068.2 D 8 0.8000 0.774 0.620 NA NA ARGFX MA1463.2 R 8 0.8000 0.754 0.603 NA NA Lhx3 MA0135.2 R 10 0.8333 0.715 0.596 NA NA Total matches= 97 (94 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 35 (10.23%) Max nb of predicted sites per peak: 8  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 8.66	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 5   oligos_6nt_mkv3_m5	k-mer sig=3.45; evalue=0.00035 csCCTCCCvs SBGGGAGGSG	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor csCCTCCCvs ZNF148 MA1653.2 D 10 1.0000 0.824 0.824 NA NA ZNF281 MA1630.3 R 10 1.0000 0.739 0.739 NA NA SP4 MA0685.2 R 9 0.9000 0.779 0.701 NA NA Total matches= 23 (20 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 35 (10.23%) Max nb of predicted sites per peak: 27  (hg38_ct_MyTrack_6202_15) Mean nb of predicted sites per peak: 11.86	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 6   oligos_6nt_mkv3_m6	k-mer sig=3.32; evalue=0.00048 taTACATAya TRTATGTATA	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor taTACATAya NFIL3 MA0025.3 R 9 0.9000 0.741 0.667 NA NA POU3F1 MA0786.2 R 9 0.8182 0.709 0.580 NA NA POU5F1B MA0792.1 R 8 0.7273 0.707 0.514 NA NA Total matches= 11 (8 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 32 (9.36%) Max nb of predicted sites per peak: 19  (hg38_ct_MyTrack_6202_1) Mean nb of predicted sites per peak: 8.53	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 7   oligos_6nt_mkv3_m7	k-mer sig=3.21; evalue=0.00062 arAAAGAAAwr YWTTTCTTTYT	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor arAAAGAAAwr STAT1::STAT2 MA0517.2 R 11 0.8462 0.708 0.599 NA NA Stat2 MA1623.2 D 9 0.7500 0.788 0.591 NA NA FOXD3 MA0041.3 D 11 0.7857 0.720 0.565 NA NA Total matches= 7 (4 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 33 (9.65%) Max nb of predicted sites per peak: 12  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 8.76	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 8   oligos_6nt_mkv3_m8	k-mer sig=2.79; evalue=0.0016 waAATAAAaa TTTTTATTTW	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor waAATAAAaa HOXB13 MA0901.3 D 9 0.9000 0.870 0.783 NA NA HOXA10 MA0899.2 D 9 0.9000 0.854 0.769 NA NA CDX4 MA1473.2 D 9 0.9000 0.829 0.746 NA NA Total matches= 43 (40 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 36 (10.53%) Max nb of predicted sites per peak: 15  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 9.58	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 9   oligos_6nt_mkv3_m9	k-mer sig=2.68; evalue=0.0021 swGACAGAGcsr YSGCTCTGTCWS	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor swGACAGAGcsr ZNF768 MA1731.2 R 8 0.6154 0.839 0.516 NA NA MEIS3 MA0775.2 D 7 0.5833 0.846 0.493 NA NA MEIS2 MA0774.1 D 8 0.6667 0.739 0.492 NA NA Total matches= 4 (1 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 31 (9.06%) Max nb of predicted sites per peak: 8  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 6.23	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 10   oligos_6nt_mkv3_m10	k-mer sig=2.41; evalue=0.0039 rrGCCaAGGcs SGCCTTGGCYY	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor rrGCCaAGGcs NFYB MA0502.3 R 9 0.8182 0.722 0.591 NA NA NFIC MA0161.3 R 7 0.6364 0.926 0.589 NA NA EBF3 MA1637.2 D 9 0.8182 0.714 0.584 NA NA Total matches= 14 (11 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 29 (8.48%) Max nb of predicted sites per peak: 6  (hg38_ct_MyTrack_6202_15) Mean nb of predicted sites per peak: 5.55	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 11   positions_6nt_m1	k-mer sig=0.75; evalue=0.18 ysAGCCAAsr YSTTGGCTSR	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor ysAGCCAAsr NFIC MA0161.3 R 7 0.7000 0.758 0.531 NA NA NFYB MA0502.3 R 8 0.7273 0.726 0.528 NA NA NFIX MA0671.2 D 6 0.6000 0.857 0.514 NA NA Total matches= 11 (8 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 31 (9.06%) Max nb of predicted sites per peak: 4  (hg38_ct_MyTrack_6202_9) Mean nb of predicted sites per peak: 5.42	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 12   positions_6nt_m2	k-mer sig=0.07; evalue=0.85 msACTTCTsy RSAGAAGTSK	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor msACTTCTsy OSR2 MA1646.2 R 8 0.8000 0.721 0.576 NA NA Nr2e1 MA0676.1 R 8 0.7273 0.707 0.514 NA NA Stat5b MA1625.2 R 6 0.4615 0.873 0.403 NA NA Total matches= 3		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 33 (9.65%) Max nb of predicted sites per peak: 6  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 4.55	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 13   oligos_7nt_mkv4_m1	k-mer sig=3.93; evalue=0.00012 haAAAATTAkm KMTAATTTTTD	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor haAAAATTAkm MEF2A MA0052.5 D 10 0.9091 0.713 0.648 NA NA BARX2 MA1471.2 D 9 0.8182 0.747 0.611 NA NA PHOX2B MA0681.3 D 10 0.7692 0.732 0.563 NA NA Total matches= 77 (74 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 36 (10.53%) Max nb of predicted sites per peak: 7  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 7.31	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 14   oligos_7nt_mkv4_m2	k-mer sig=2.67; evalue=0.0021 csAGGCtGGrs SYCCAGCCTSG	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor csAGGCtGGrs FEZF2 MA2341.1 R 8 0.7273 0.940 0.683 NA NA ZNF213 MA2121.1 D 10 0.7692 0.770 0.593 NA NA ZNF770 MA2099.1 R 8 0.7273 0.801 0.583 NA NA Total matches= 8 (5 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 33 (9.65%) Max nb of predicted sites per peak: 8  (hg38_ct_MyTrack_6202_9) Mean nb of predicted sites per peak: 9.67	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 15   oligos_7nt_mkv4_m3	k-mer sig=1.61; evalue=0.025 maAAAACAAaa TTTTGTTTTTK	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor maAAAACAAaa Foxq1 MA0040.2 R 10 0.9091 0.730 0.664 NA NA Foxj3 MA0851.2 D 9 0.8182 0.746 0.610 NA NA Sox6 MA0515.1 R 9 0.7500 0.785 0.589 NA NA Total matches= 38 (35 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 33 (9.65%) Max nb of predicted sites per peak: 6  (hg38_ct_MyTrack_6202_4) Mean nb of predicted sites per peak: 7.27	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 16   oligos_7nt_mkv4_m4	k-mer sig=1.51; evalue=0.031 rtAAGGCAAta TATTGCCTTAY	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor rtAAGGCAAta Nr2e1 MA0676.1 D 9 0.8182 0.765 0.626 NA NA Zfp335 MA2002.2 D 7 0.6364 0.768 0.489 NA NA TFAP2B MA0811.2 D 8 0.5714 0.751 0.429 NA NA Total matches= 5 (2 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 26 (7.60%) Max nb of predicted sites per peak: 4  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 2.85	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 17   oligos_7nt_mkv4_m5	k-mer sig=1.08; evalue=0.083 vhCAGCATCwy RWGATGCTGDB	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor vhCAGCATCwy MAFF MA0495.4 D 11 1.0000 0.724 0.724 NA NA Mafb MA0117.3 R 11 1.0000 0.707 0.707 NA NA ZNF549 MA1728.2 R 7 0.5833 0.790 0.461 NA NA Total matches= 11 (8 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 26 (7.60%) Max nb of predicted sites per peak: 3  (hg38_ct_MyTrack_6202_21) Mean nb of predicted sites per peak: 3.31	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 18   oligos_7nt_mkv4_m6	k-mer sig=1.03; evalue=0.093 cwGGGATTAya TRTAATCCCWG	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor cwGGGATTAya PITX2 MA1547.2 R 8 0.7273 0.934 0.679 NA NA Dmbx1 MA0883.2 D 9 0.7500 0.859 0.644 NA NA OTX2 MA0712.3 D 7 0.6364 0.959 0.611 NA NA Total matches= 18 (15 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 34 (9.94%) Max nb of predicted sites per peak: 7  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 6.32	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 19   oligos_7nt_mkv4_m7	k-mer sig=1.03; evalue=0.093 smGCCtGGCCmv BKGGCCAGGCKS	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor smGCCtGGCCmv ZNF213 MA2121.1 R 10 0.7143 0.829 0.592 NA NA Zfp335 MA2002.2 R 7 0.5833 0.761 0.444 NA NA Plagl1 MA1615.2 R 7 0.5385 0.819 0.441 NA NA Total matches= 4 (1 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 31 (9.06%) Max nb of predicted sites per peak: 32  (hg38_ct_MyTrack_6202_15) Mean nb of predicted sites per peak: 11.74	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 20   oligos_7nt_mkv4_m8	k-mer sig=0.46; evalue=0.35 dwAATGCAAwm KWTTGCATTWH	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor dwAATGCAAwm CEBPG MA1636.2 D 10 0.9091 0.735 0.668 NA NA POU3F1 MA0786.2 D 10 0.9091 0.734 0.667 NA NA ATF4 MA0833.3 D 10 0.9091 0.730 0.664 NA NA Total matches= 22 (19 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 29 (8.48%) Max nb of predicted sites per peak: 3  (hg38_ct_MyTrack_6202_5) Mean nb of predicted sites per peak: 2.93	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 21   oligos_7nt_mkv4_m9	k-mer sig=0.21; evalue=0.62 adGAAAAGAAaa TTTTCTTTTCHT	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor adGAAAAGAAaa ZIM3 MA1709.2 D 10 0.7692 0.712 0.548 NA NA ZNF677 MA2101.1 D 10 0.7143 0.741 0.529 NA NA ZNF384 MA1125.2 D 8 0.6667 0.793 0.529 NA NA Total matches= 8 (5 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 24 (7.02%) Max nb of predicted sites per peak: 6  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 3.50	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 22   oligos_7nt_mkv4_m10	k-mer sig=0.07; evalue=0.85 raAAAGGAAdr YHTTCCTTTTY	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor raAAAGGAAdr FOXJ2::ELF1 MA1952.2 D 11 1.0000 0.724 0.724 NA NA ETV2::FOXI1 MA1942.2 D 11 1.0000 0.710 0.710 NA NA ERF::FOXI1 MA1935.2 D 10 0.9091 0.752 0.684 NA NA Total matches= 34 (31 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 28 (8.19%) Max nb of predicted sites per peak: 9  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 6.04	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 23   oligos_8nt_mkv5_m1	k-mer sig=3.78; evalue=0.00017 cbCCtCCtCCys SRGGAGGAGGVG	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor cbCCtCCtCCys Wt1 MA1627.2 D 10 0.8333 0.763 0.636 NA NA ZNF281 MA1630.3 R 10 0.8333 0.761 0.635 NA NA ZNF148 MA1653.2 D 10 0.8333 0.742 0.619 NA NA Total matches= 11 (8 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 31 (9.06%) Max nb of predicted sites per peak: 26  (hg38_ct_MyTrack_6202_9) Mean nb of predicted sites per peak: 9.32	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 24   oligos_8nt_mkv5_m2	k-mer sig=1.77; evalue=0.017 awACAAAAACwa TWGTTTTTGTWT	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor awACAAAAACwa ZNF384 MA1125.2 D 8 0.6667 0.826 0.551 NA NA Stat2 MA1623.2 D 9 0.6923 0.740 0.513 NA NA SOX4 MA0867.3 D 8 0.6667 0.721 0.481 NA NA Total matches= 13 (10 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 32 (9.36%) Max nb of predicted sites per peak: 6  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 4.91	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 25   dyads_m1	k-mer sig=39.36; evalue=4.4e-40 twtattTaTTTTAaww WWTTAAAATAAATAWA	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor twtattTaTTTTAaww FOXD3 MA0041.3 R 14 0.8750 0.708 0.620 NA NA ONECUT3 MA0757.2 R 12 0.7500 0.723 0.542 NA NA FOXD2 MA0847.4 R 11 0.6875 0.727 0.500 NA NA Total matches= 11 (8 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 35 (10.23%) Max nb of predicted sites per peak: 17  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 8.57	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 26   dyads_m2	k-mer sig=32.32; evalue=4.8e-33 csCCyCCCCCCCCCCmSCCCyCss SSGRGGGSKGGGGGGGGGGRGGSG	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor csCCyCCCCCCCCCCmSCCCyCss PRDM9 MA1723.2 R 16 0.5714 0.751 0.429 NA NA Total matches= 1		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 30 (8.77%) Max nb of predicted sites per peak: 414  (hg38_ct_MyTrack_6202_15) Mean nb of predicted sites per peak: 52.43	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 27   dyads_m3	k-mer sig=35.65; evalue=2.2e-36 wwttaAAAATAaatawa TWTATTTATTTTTAAWW	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor wwttaAAAATAaatawa FOXD3 MA0041.3 D 14 0.8235 0.711 0.586 NA NA ONECUT3 MA0757.2 D 12 0.7059 0.795 0.561 NA NA MEF2C MA0497.2 D 11 0.6471 0.822 0.532 NA NA Total matches= 8 (5 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 36 (10.53%) Max nb of predicted sites per peak: 20  (hg38_ct_MyTrack_6202_8) Mean nb of predicted sites per peak: 12.22	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 28   dyads_m4	k-mer sig=26.31; evalue=4.9e-27 srGCCtGGCys SRGCCAGGCYS	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor srGCCtGGCys ZNF213 MA2121.1 R 10 0.7692 0.791 0.609 NA NA Hand1::Tcf3 MA0092.2 D 9 0.8182 0.704 0.576 NA NA Zfp335 MA2002.2 R 7 0.6364 0.768 0.489 NA NA Total matches= 6 (3 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 31 (9.06%) Max nb of predicted sites per peak: 13  (hg38_ct_MyTrack_6202_15) Mean nb of predicted sites per peak: 9.74	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 29   dyads_m5	k-mer sig=20.43; evalue=3.7e-21 ssCCCAGGCys SRGCCTGGGSS	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor ssCCCAGGCys ZNF213 MA2121.1 D 10 0.7692 0.952 0.732 NA NA Zfp809 MA2125.1 D 9 0.8182 0.761 0.623 NA NA FEZF2 MA2341.1 D 8 0.7273 0.796 0.579 NA NA Total matches= 10 (7 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 32 (9.36%) Max nb of predicted sites per peak: 10  (hg38_ct_MyTrack_6202_15) Mean nb of predicted sites per peak: 6.69	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 30   dyads_m6	k-mer sig=19.21; evalue=6.2e-20 csCCCaCCss SSGGTGGGSG	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor csCCCaCCss KLF5 MA0599.1 D 10 1.0000 0.781 0.781 NA NA PATZ1 MA1961.2 R 10 0.9091 0.800 0.727 NA NA ZBTB7B MA0694.2 D 10 1.0000 0.705 0.705 NA NA Total matches= 37 (34 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 33 (9.65%) Max nb of predicted sites per peak: 33  (hg38_ct_MyTrack_6202_15) Mean nb of predicted sites per peak: 10.03	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 31   dyads_m7	k-mer sig=18.62; evalue=2.4e-19 ssCCCCryCTCys SRGAGRYGGGGSS	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor ssCCCCryCTCys PATZ1 MA1961.2 R 11 0.8462 0.750 0.635 NA NA ZNF281 MA1630.3 R 10 0.7692 0.702 0.540 NA NA SP2 MA0516.3 R 9 0.6923 0.719 0.498 NA NA Total matches= 8 (5 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 31 (9.06%) Max nb of predicted sites per peak: 35  (hg38_ct_MyTrack_6202_15) Mean nb of predicted sites per peak: 11.10	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 32   dyads_m8	k-mer sig=27.42; evalue=3.8e-28 wwtTAAAAAgaaAwaatww WWATTWTTTCTTTTTAAWW	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor wwtTAAAAAgaaAwaatww ONECUT3 MA0757.2 D 12 0.6316 0.738 0.466 NA NA Total matches= 1		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 33 (9.65%) Max nb of predicted sites per peak: 14  (hg38_ct_MyTrack_6202_0) Mean nb of predicted sites per peak: 10.88	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 33   dyads_m9	k-mer sig=18.62; evalue=2.4e-19 CcGCCCCCCtCCSCCtCCCyCms SKGRGGGAGGSGGAGGGGGGCGG	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor CcGCCCCCCtCCSCCtCCCyCms ZNF460 MA1596.1 D 16 0.6957 0.777 0.540 NA NA PATZ1 MA1961.2 R 11 0.4783 0.883 0.423 NA NA ZNF281 MA1630.3 R 10 0.4348 0.961 0.418 NA NA Total matches= 4 (1 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 32 (9.36%) Max nb of predicted sites per peak: 223  (hg38_ct_MyTrack_6202_15) Mean nb of predicted sites per peak: 31.59	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]
Motif 34   dyads_m10	k-mer sig=13.89; evalue=1.3e-14 ssCGCsssGCCCGss SSCGGGCSSSGCGSS	[ k-mers: statistics assemblies ] [ matrix: tab_format transfac_format ]
JASPAR2024_CORE_vertebrates_non-redundant name id strand Nb overlap % Pearson Normalized aligned col. motif aligned col. match       columns aligned correlation cor ssCGCsssGCCCGss ZBED4 MA2328.1 D 10 0.6667 0.851 0.567 NA NA SP3 MA0746.3 D 11 0.7333 0.722 0.529 NA NA KLF3 MA1516.2 D 10 0.6667 0.738 0.492 NA NA Total matches= 15 (12 more)		[ match table: html text ] [ alignments (logos): html text ]
Predicted sites on test peaks	Distribution of sites Nb of predicted sites per peak Nb peaks with at least one predicted site: 32 (9.36%) Max nb of predicted sites per peak: 216  (hg38_ct_MyTrack_6202_15) Mean nb of predicted sites per peak: 24.50	[ sites: text ] [ distribution: text ] [ sites per peak: text ] [ enrichment: text graph ]

Command:  peak-motifs  -v 1 -title SMAD2_helasherg_SEACR_stringent_peak_motifs -i $RSAT/public_html/tmp/www-data/2026/03/19/peak-motifs.2026-03-19.143247_2026-03-19.143247_dTipSp/peak-motifspeak_seq -max_seq_len 1000 -markov auto -disco oligos,dyads,positions,local_words -nmotifs 10 -minol 6 -maxol 8 -no_merge_lengths -2str -origin center -motif_db JASPAR2024_CORE_vertebrates_non-redundant tf $RSAT/public_html/motif_databases/JASPAR/Jaspar_2024/JASPAR2024_CORE_vertebrates_non-redundant_pfms_transfac.txt -scan_markov 1 -task purge,seqlen,composition,disco,merge_motifs,split_motifs,motifs_vs_motifs,timelog,archive,synthesis,small_summary,motifs_vs_db,scan -prefix peak-motifs -noov -img_format png -outdir $RSAT/public_html/tmp/www-data/2026/03/19/peak-motifs.2026-03-19.143247_2026-03-19.143247_dTipSp

Warning: Not a single pattern was found in file tmp/www-data/2026/03/19/peak-motifs.2026-03-19.143247_2026-03-19.143247_dTipSp/results/positions_7nt/peak-motifs_positions-2str-noov_7nt_ci50.tab

Warning: Not a single pattern was found in file tmp/www-data/2026/03/19/peak-motifs.2026-03-19.143247_2026-03-19.143247_dTipSp/results/positions_8nt/peak-motifs_positions-2str-noov_8nt_ci50.tab

Warning: Not a single pattern was found in file tmp/www-data/2026/03/19/peak-motifs.2026-03-19.143247_2026-03-19.143247_dTipSp/results/local_words_6nt/peak-motifs_local_words-2str-noov_6nt_windgroup50.tab

Warning: Not a single pattern was found in file tmp/www-data/2026/03/19/peak-motifs.2026-03-19.143247_2026-03-19.143247_dTipSp/results/local_words_7nt/peak-motifs_local_words-2str-noov_7nt_windgroup50.tab

Warning: Not a single pattern was found in file tmp/www-data/2026/03/19/peak-motifs.2026-03-19.143247_2026-03-19.143247_dTipSp/results/local_words_8nt/peak-motifs_local_words-2str-noov_8nt_windgroup50.tab