position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/03/30/peak-motifs.2026-03-30.164132_2026-03-30.164132_qGS9No/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title aniger -origin center -offset 6 -o $RSAT/public_html/tmp/www-data/2026/03/30/peak-motifs.2026-03-30.164132_2026-03-30.164132_qGS9No/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/03/30/peak-motifs.2026-03-30.164132_2026-03-30.164132_qGS9No/data/sequences/peak-motifs_test_maxlen1000_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/03/30/peak-motifs.2026-03-30.164132_2026-03-30.164132_qGS9No/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 37 Sum of sequence lengths 32964 Min sequence length 0 Max sequence length 1000 Average sequence length 890 Possible positions 32927 Sequences: # length ID 1 1000 pid_1787_+ 2 1000 pid_4473_- 3 1000 pid_10881_+ 4 1000 pid_9810_+ 5 955 pid_11670_- 6 1000 pid_10052_- 7 1000 pid_4471_+ 8 1000 pid_8708_+ 9 1000 pid_9019_- 10 1000 pid_8994_+ 11 1000 pid_3339_- 12 1000 pid_8911_- 13 591 pid_1069_+ 14 1000 pid_9001_- 15 1000 pid_9251_- 16 1000 pid_1918_- 17 835 pid_11715_- 18 320 pid_2451_+ 19 1000 pid_1071_+ 20 1000 pid_7_- 21 1000 pid_819_- 22 630 pid_9556_+ 23 543 pid_1952_- 24 1000 pid_10868_+ 25 1000 pid_8707_- 26 448 pid_11685_+ 27 1000 pid_2929_+ 28 1000 pid_9002_+ 29 1000 pid_268_+ 30 841 pid_10870_+ 31 495 pid_7382_+ 32 1000 pid_8993_- 33 1000 pid_3928_- 34 593 pid_4917_+ 35 1000 pid_1648_- 36 713 pid_4328_- 37 1000 pid_4034_+ Oligonucleotide statistics: Total occurrences 31897 Position interval parameters: Position interval 20 Number of windows 51 Total positions 32927 Degrees of freedom 50 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -26 [-519,-500] -509.5 26 156 2 -25 [-499,-480] -489.5 27 524 3 -24 [-479,-460] -469.5 27 540 4 -23 [-459,-440] -449.5 27 540 5 -22 [-439,-420] -429.5 29 551 6 -21 [-419,-400] -409.5 29 580 7 -20 [-399,-380] -389.5 29 580 8 -19 [-379,-360] -369.5 30 583 9 -18 [-359,-340] -349.5 30 600 10 -17 [-339,-320] -329.5 31 602 11 -16 [-319,-300] -309.5 33 625 12 -15 [-299,-280] -289.5 33 660 13 -14 [-279,-260] -269.5 34 678 14 -13 [-259,-240] -249.5 35 694 15 -12 [-239,-220] -229.5 36 710 16 -11 [-219,-200] -209.5 36 720 17 -10 [-199,-180] -189.5 36 720 18 -9 [-179,-160] -169.5 37 726 19 -8 [-159,-140] -149.5 37 740 20 -7 [-139,-120] -129.5 37 740 21 -6 [-119,-100] -109.5 37 740 22 -5 [-99,-80] -89.5 37 740 23 -4 [-79,-60] -69.5 37 740 24 -3 [-59,-40] -49.5 37 740 25 -2 [-39,-20] -29.5 37 740 26 -1 [-19,0] -9.5 37 740 27 0 [1,20] 10.5 37 740 28 1 [21,40] 30.5 37 740 29 2 [41,60] 50.5 37 740 30 3 [61,80] 70.5 37 740 31 4 [81,100] 90.5 37 740 32 5 [101,120] 110.5 37 740 33 6 [121,140] 130.5 37 740 34 7 [141,160] 150.5 37 733 35 8 [161,180] 170.5 36 720 36 9 [181,200] 190.5 36 720 37 10 [201,220] 210.5 36 717 38 11 [221,240] 230.5 35 700 39 12 [241,260] 250.5 34 680 40 13 [261,280] 270.5 34 664 41 14 [281,300] 290.5 33 637 42 15 [301,320] 310.5 31 607 43 16 [321,340] 330.5 30 600 44 17 [341,360] 350.5 30 589 45 18 [361,380] 370.5 29 580 46 19 [381,400] 390.5 29 580 47 20 [401,420] 410.5 29 563 48 21 [421,440] 430.5 27 540 49 22 [441,460] 450.5 27 540 50 23 [461,480] 470.5 27 530 51 24 [481,500] 490.5 26 338 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -509.5 occurrences in window 1 [-519,-500] 11 -489.5 occurrences in window 2 [-499,-480] 12 -469.5 occurrences in window 3 [-479,-460] 13 -449.5 occurrences in window 4 [-459,-440] 14 -429.5 occurrences in window 5 [-439,-420] 15 -409.5 occurrences in window 6 [-419,-400] 16 -389.5 occurrences in window 7 [-399,-380] 17 -369.5 occurrences in window 8 [-379,-360] 18 -349.5 occurrences in window 9 [-359,-340] 19 -329.5 occurrences in window 10 [-339,-320] 20 -309.5 occurrences in window 11 [-319,-300] 21 -289.5 occurrences in window 12 [-299,-280] 22 -269.5 occurrences in window 13 [-279,-260] 23 -249.5 occurrences in window 14 [-259,-240] 24 -229.5 occurrences in window 15 [-239,-220] 25 -209.5 occurrences in window 16 [-219,-200] 26 -189.5 occurrences in window 17 [-199,-180] 27 -169.5 occurrences in window 18 [-179,-160] 28 -149.5 occurrences in window 19 [-159,-140] 29 -129.5 occurrences in window 20 [-139,-120] 30 -109.5 occurrences in window 21 [-119,-100] 31 -89.5 occurrences in window 22 [-99,-80] 32 -69.5 occurrences in window 23 [-79,-60] 33 -49.5 occurrences in window 24 [-59,-40] 34 -29.5 occurrences in window 25 [-39,-20] 35 -9.5 occurrences in window 26 [-19,0] 36 10.5 occurrences in window 27 [1,20] 37 30.5 occurrences in window 28 [21,40] 38 50.5 occurrences in window 29 [41,60] 39 70.5 occurrences in window 30 [61,80] 40 90.5 occurrences in window 31 [81,100] 41 110.5 occurrences in window 32 [101,120] 42 130.5 occurrences in window 33 [121,140] 43 150.5 occurrences in window 34 [141,160] 44 170.5 occurrences in window 35 [161,180] 45 190.5 occurrences in window 36 [181,200] 46 210.5 occurrences in window 37 [201,220] 47 230.5 occurrences in window 38 [221,240] 48 250.5 occurrences in window 39 [241,260] 49 270.5 occurrences in window 40 [261,280] 50 290.5 occurrences in window 41 [281,300] 51 310.5 occurrences in window 42 [301,320] 52 330.5 occurrences in window 43 [321,340] 53 350.5 occurrences in window 44 [341,360] 54 370.5 occurrences in window 45 [361,380] 55 390.5 occurrences in window 46 [381,400] 56 410.5 occurrences in window 47 [401,420] 57 430.5 occurrences in window 48 [421,440] 58 450.5 occurrences in window 49 [441,460] 59 470.5 occurrences in window 50 [461,480] 60 490.5 occurrences in window 51 [481,500]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -509.5 | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| tc | tc | 2437 | 100.5 | 50 | 3.0e-05 | 0.00049 | 3.31 | 1 | 14 | 25 | 32 | 27 | 28 | 34 | 46 | 38 | 41 | 29 | 44 | 45 | 42 | 51 | 43 | 40 | 48 | 47 | 63 | 48 | 43 | 53 | 62 | 58 | 51 | 52 | 43 | 56 | 48 | 56 | 53 | 47 | 66 | 62 | 54 | 56 | 63 | 53 | 50 | 52 | 46 | 63 | 50 | 51 | 46 | 46 | 69 | 58 | 57 | 51 | 37 |
| aa | aa | 2130 | 94.2 | 50 | 1.6e-04 | 0.0025 | 2.59 | 2 | 7 | 32 | 22 | 30 | 19 | 31 | 35 | 22 | 43 | 40 | 48 | 41 | 47 | 41 | 52 | 56 | 50 | 54 | 30 | 55 | 52 | 48 | 66 | 45 | 64 | 45 | 64 | 48 | 56 | 54 | 52 | 40 | 30 | 42 | 52 | 53 | 40 | 51 | 32 | 38 | 34 | 39 | 35 | 32 | 43 | 45 | 31 | 26 | 38 | 55 | 25 |
| tt | tt | 2340 | 88.5 | 50 | 6.5e-04 | 0.01 | 1.98 | 3 | 10 | 38 | 45 | 41 | 52 | 38 | 39 | 38 | 54 | 33 | 38 | 37 | 46 | 50 | 38 | 48 | 47 | 41 | 66 | 49 | 54 | 49 | 36 | 56 | 44 | 53 | 54 | 47 | 49 | 52 | 44 | 36 | 56 | 50 | 49 | 56 | 49 | 44 | 73 | 37 | 51 | 47 | 44 | 34 | 57 | 44 | 52 | 49 | 63 | 44 | 19 |
| gg | gg | 1687 | 81.4 | 50 | 3.3e-03 | 0.053 | 1.28 | 4 | 6 | 24 | 23 | 34 | 31 | 36 | 27 | 28 | 21 | 32 | 31 | 33 | 38 | 32 | 51 | 40 | 43 | 40 | 48 | 40 | 36 | 40 | 25 | 29 | 32 | 38 | 29 | 36 | 46 | 38 | 35 | 53 | 44 | 43 | 37 | 39 | 37 | 54 | 39 | 46 | 23 | 29 | 35 | 37 | 24 | 27 | 14 | 14 | 21 | 21 | 8 |
| ca | ca | 2202 | 80.9 | 50 | 3.7e-03 | 0.06 | 1.23 | 5 | 11 | 26 | 22 | 33 | 25 | 34 | 32 | 30 | 35 | 41 | 39 | 44 | 47 | 48 | 42 | 49 | 44 | 54 | 55 | 45 | 48 | 45 | 61 | 59 | 51 | 47 | 45 | 55 | 44 | 48 | 52 | 60 | 42 | 45 | 54 | 47 | 40 | 44 | 34 | 50 | 42 | 46 | 48 | 42 | 37 | 38 | 38 | 42 | 49 | 42 | 51 |
| ct | ct | 2291 | 70.2 | 50 | 3.1e-02 | 0.5 | 0.30 | 6 | 9 | 32 | 37 | 33 | 32 | 32 | 42 | 38 | 35 | 31 | 38 | 49 | 46 | 41 | 52 | 45 | 47 | 47 | 56 | 42 | 39 | 47 | 57 | 52 | 47 | 52 | 54 | 46 | 42 | 49 | 52 | 46 | 54 | 60 | 56 | 46 | 64 | 49 | 46 | 47 | 47 | 52 | 40 | 51 | 34 | 54 | 64 | 57 | 47 | 41 | 15 |
| cc | cc | 2375 | 57.7 | 50 | 2.1e-01 | 3.4 | -0.53 | 7 | 9 | 24 | 30 | 34 | 36 | 34 | 35 | 39 | 37 | 46 | 36 | 44 | 46 | 50 | 46 | 44 | 40 | 56 | 56 | 55 | 51 | 47 | 50 | 59 | 52 | 66 | 58 | 55 | 51 | 58 | 56 | 55 | 59 | 47 | 57 | 61 | 50 | 43 | 56 | 45 | 59 | 55 | 39 | 38 | 53 | 36 | 49 | 58 | 51 | 42 | 22 |
| ta | ta | 1794 | 57.2 | 50 | 2.2e-01 | 3.6 | -0.56 | 8 | 7 | 32 | 29 | 28 | 26 | 26 | 21 | 22 | 32 | 30 | 43 | 49 | 37 | 54 | 44 | 39 | 39 | 46 | 34 | 42 | 43 | 42 | 30 | 38 | 48 | 40 | 50 | 37 | 37 | 33 | 36 | 43 | 35 | 42 | 45 | 39 | 36 | 30 | 37 | 33 | 39 | 32 | 32 | 32 | 35 | 52 | 28 | 28 | 20 | 26 | 16 |
| at | at | 2356 | 56.2 | 50 | 2.5e-01 | 4.1 | -0.61 | 9 | 14 | 33 | 34 | 27 | 35 | 36 | 27 | 39 | 54 | 41 | 49 | 50 | 40 | 68 | 33 | 52 | 57 | 50 | 51 | 56 | 55 | 53 | 58 | 55 | 61 | 44 | 46 | 61 | 67 | 46 | 45 | 59 | 49 | 45 | 56 | 58 | 46 | 51 | 55 | 44 | 46 | 51 | 49 | 41 | 47 | 50 | 36 | 34 | 31 | 40 | 31 |
| ga | ga | 1851 | 54.8 | 50 | 3.0e-01 | 4.8 | -0.68 | 10 | 8 | 34 | 30 | 31 | 31 | 37 | 30 | 37 | 38 | 35 | 36 | 31 | 40 | 37 | 43 | 38 | 40 | 40 | 38 | 50 | 55 | 48 | 44 | 38 | 38 | 50 | 45 | 43 | 49 | 37 | 40 | 34 | 39 | 51 | 38 | 43 | 44 | 54 | 41 | 46 | 30 | 22 | 36 | 33 | 32 | 22 | 17 | 21 | 24 | 24 | 9 |
| ac | ac | 1724 | 52.1 | 50 | 3.9e-01 | 6.3 | -0.80 | 11 | 4 | 22 | 23 | 25 | 20 | 29 | 28 | 16 | 23 | 26 | 37 | 37 | 38 | 34 | 45 | 39 | 35 | 41 | 39 | 41 | 49 | 35 | 47 | 39 | 44 | 49 | 41 | 47 | 31 | 32 | 44 | 36 | 28 | 47 | 41 | 36 | 37 | 32 | 24 | 40 | 30 | 32 | 34 | 33 | 30 | 34 | 26 | 32 | 34 | 34 | 24 |
| gc | gc | 1813 | 50.6 | 50 | 4.5e-01 | 7.2 | -0.86 | 12 | 10 | 33 | 30 | 26 | 32 | 27 | 30 | 42 | 27 | 39 | 23 | 34 | 42 | 34 | 35 | 46 | 49 | 43 | 40 | 32 | 32 | 41 | 42 | 41 | 36 | 38 | 36 | 41 | 43 | 49 | 42 | 56 | 47 | 35 | 45 | 34 | 46 | 37 | 45 | 40 | 45 | 26 | 37 | 35 | 23 | 28 | 29 | 26 | 19 | 22 | 23 |
| ag | ag | 1766 | 47.3 | 50 | 5.8e-01 | 9.3 | -0.97 | 13 | 7 | 38 | 29 | 34 | 29 | 33 | 27 | 35 | 29 | 35 | 32 | 41 | 41 | 39 | 42 | 40 | 35 | 49 | 31 | 48 | 38 | 44 | 37 | 39 | 31 | 43 | 51 | 31 | 32 | 42 | 36 | 50 | 31 | 45 | 38 | 39 | 35 | 47 | 37 | 37 | 40 | 18 | 31 | 32 | 28 | 29 | 23 | 21 | 22 | 25 | 20 |
| tg | tg | 2043 | 46.8 | 50 | 6.0e-01 | 9.6 | -0.98 | 14 | 8 | 30 | 37 | 33 | 46 | 38 | 36 | 47 | 34 | 47 | 35 | 37 | 45 | 35 | 45 | 51 | 59 | 36 | 43 | 43 | 51 | 48 | 48 | 45 | 46 | 42 | 40 | 48 | 51 | 48 | 56 | 40 | 52 | 38 | 34 | 45 | 50 | 53 | 42 | 41 | 40 | 39 | 38 | 34 | 34 | 31 | 35 | 31 | 25 | 24 | 9 |
| cg | cg | 1470 | 37.5 | 50 | 9.0e-01 | 14 | -1.16 | 15 | 7 | 22 | 24 | 15 | 23 | 25 | 29 | 25 | 21 | 24 | 23 | 26 | 28 | 28 | 34 | 34 | 32 | 30 | 36 | 35 | 35 | 39 | 29 | 30 | 36 | 33 | 26 | 38 | 39 | 39 | 33 | 35 | 44 | 36 | 37 | 30 | 42 | 29 | 36 | 36 | 33 | 23 | 26 | 34 | 24 | 17 | 20 | 20 | 17 | 18 | 15 |
| gt | gt | 1618 | 31.1 | 50 | 9.8e-01 | 16 | -1.20 | 16 | 7 | 19 | 33 | 29 | 26 | 30 | 36 | 27 | 22 | 33 | 33 | 34 | 35 | 32 | 45 | 39 | 35 | 32 | 34 | 39 | 39 | 41 | 28 | 37 | 39 | 28 | 38 | 31 | 35 | 39 | 44 | 30 | 44 | 35 | 27 | 38 | 38 | 29 | 33 | 32 | 32 | 33 | 26 | 30 | 33 | 27 | 32 | 23 | 22 | 21 | 14 |
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