position-analysis -v 1 -i $RSAT/public_html/tmp/www-data/2026/05/27/peak-motifs.2026-05-27.215012_2026-05-27.215012_WiGkpW/data/sequences/peak-motifs_test_maxlen1200_purged_ml40_mis3.fasta -format fasta -sort -return html,chi,sig,distrib,graphs,rank,index -max_graphs 20 -1str -ovlp -seqtype dna -l 2 -ci 20 -img_format png -title ld -origin center -offset 0 -o $RSAT/public_html/tmp/www-data/2026/05/27/peak-motifs.2026-05-27.215012_2026-05-27.215012_WiGkpW/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Citation: van Helden, et al. (2000). Nucleic Acids Res 28, 1000-1010. Sequence file $RSAT/public_html/tmp/www-data/2026/05/27/peak-motifs.2026-05-27.215012_2026-05-27.215012_WiGkpW/data/sequences/peak-motifs_test_maxlen1200_purged_ml40_mis3.fasta Sequence format fasta Sequence type dna Output file $RSAT/public_html/tmp/www-data/2026/05/27/peak-motifs.2026-05-27.215012_2026-05-27.215012_WiGkpW/results/composition/peak-motifs_test_profiles-1str-ovlp_2nt_ci20.tab Oligo length 2 Occurrences counted on a single strands Conditions of applicability checked. Background model estimation: homogeneous repartition Sequence statistics: Nb of sequences 33 Sum of sequence lengths 18937 Min sequence length 0 Max sequence length 1088 Average sequence length 573 Possible positions 18904 Sequences: # length ID 1 939 FBgn0001248|Idh 2 893 FBgn0002719|Men 3 284 FBgn0010548|Aldh-III 4 431 FBgn0012034|AcCoAS 5 698 FBgn0012036|Aldh 6 699 FBgn0015623|Cpr 7 897 FBgn0020236|ATPCL 8 363 FBgn0025352|Mtpbeta 9 445 FBgn0025814|Mgstl 10 137 FBgn0028479|Mtpalpha 11 983 FBgn0028646|aralar1 12 230 FBgn0030607|dob 13 727 FBgn0030608|Lsd-2 14 423 FBgn0031069|Abcd3 15 456 FBgn0032265|CG18301 16 486 FBgn0033246|ACC 17 199 FBgn0033451|Marc 18 148 FBgn0035770|pst 19 706 FBgn0036449|bmm 20 906 FBgn0036821|CG3961 21 326 FBgn0037265|spartin 22 885 FBgn0037607|CG8036 23 486 FBgn0039156|CG6178 24 213 FBgn0039543|CROT 25 191 FBgn0039697|Etfb 26 518 FBgn0040064|yip2 27 231 FBgn0040336|Seipin 28 1088 FBgn0086687|Desat1 29 1082 FBgn0262782|Mdh1 30 974 FBgn0263120|Acsl 31 975 FBgn0264294|Cyt-b5 32 751 FBgn0283427|FASN1 33 167 FBgn0286783|Etf-QO Oligonucleotide statistics: Total occurrences 18862 Position interval parameters: Position interval 20 Number of windows 56 Total positions 18904 Degrees of freedom 55 K-mer clustering parameters: Number of clusters 0 Clustering method complete Position intervals: window [min,max] mid seq occ 1 -28 [-559,-540] -549.5 2 6 2 -27 [-539,-520] -529.5 2 40 3 -26 [-519,-500] -509.5 2 40 4 -25 [-499,-480] -489.5 5 68 5 -24 [-479,-460] -469.5 6 110 6 -23 [-459,-440] -449.5 10 153 7 -22 [-439,-420] -429.5 10 200 8 -21 [-419,-400] -409.5 10 200 9 -20 [-399,-380] -389.5 10 200 10 -19 [-379,-360] -369.5 12 220 11 -18 [-359,-340] -349.5 15 274 12 -17 [-339,-320] -329.5 15 300 13 -16 [-319,-300] -309.5 15 300 14 -15 [-299,-280] -289.5 15 300 15 -14 [-279,-260] -269.5 15 300 16 -13 [-259,-240] -249.5 18 328 17 -12 [-239,-220] -229.5 20 371 18 -11 [-219,-200] -209.5 22 428 19 -10 [-199,-180] -189.5 23 442 20 -9 [-179,-160] -169.5 24 464 21 -8 [-159,-140] -149.5 25 482 22 -7 [-139,-120] -129.5 25 500 23 -6 [-119,-100] -109.5 28 539 24 -5 [-99,-80] -89.5 31 600 25 -4 [-79,-60] -69.5 33 643 26 -3 [-59,-40] -49.5 33 660 27 -2 [-39,-20] -29.5 33 660 28 -1 [-19,0] -9.5 33 660 29 0 [1,20] 10.5 33 660 30 1 [21,40] 30.5 33 660 31 2 [41,60] 50.5 33 660 32 3 [61,80] 70.5 33 640 33 4 [81,100] 90.5 31 594 34 5 [101,120] 110.5 28 532 35 6 [121,140] 130.5 25 500 36 7 [141,160] 150.5 25 481 37 8 [161,180] 170.5 24 461 38 9 [181,200] 190.5 22 440 39 10 [201,220] 210.5 22 424 40 11 [221,240] 230.5 20 368 41 12 [241,260] 250.5 18 319 42 13 [261,280] 270.5 15 300 43 14 [281,300] 290.5 15 300 44 15 [301,320] 310.5 15 300 45 16 [321,340] 330.5 15 300 46 17 [341,360] 350.5 15 266 47 18 [361,380] 370.5 12 216 48 19 [381,400] 390.5 10 200 49 20 [401,420] 410.5 10 200 50 21 [421,440] 430.5 10 200 51 22 [441,460] 450.5 10 144 52 23 [461,480] 470.5 6 108 53 24 [481,500] 490.5 5 61 54 25 [501,520] 510.5 2 40 55 26 [521,540] 530.5 2 39 56 27 [541,560] 550.5 1 3 Column headers 1 seq pattern sequence 2 id pattern identifier 3 occ pattern occurrences 4 chi2 observed chi-square 5 df degrees of freedom 6 Pval P-value (probability for one word to be a false positive) 7 Eval E-value; expected number of false positives (Eval = Pval * nb_tests) 8 sig Significance (sig = -log10(Eval)) 9 rank rank of the pattern according to sorting criterion 10 -549.5 occurrences in window 1 [-559,-540] 11 -529.5 occurrences in window 2 [-539,-520] 12 -509.5 occurrences in window 3 [-519,-500] 13 -489.5 occurrences in window 4 [-499,-480] 14 -469.5 occurrences in window 5 [-479,-460] 15 -449.5 occurrences in window 6 [-459,-440] 16 -429.5 occurrences in window 7 [-439,-420] 17 -409.5 occurrences in window 8 [-419,-400] 18 -389.5 occurrences in window 9 [-399,-380] 19 -369.5 occurrences in window 10 [-379,-360] 20 -349.5 occurrences in window 11 [-359,-340] 21 -329.5 occurrences in window 12 [-339,-320] 22 -309.5 occurrences in window 13 [-319,-300] 23 -289.5 occurrences in window 14 [-299,-280] 24 -269.5 occurrences in window 15 [-279,-260] 25 -249.5 occurrences in window 16 [-259,-240] 26 -229.5 occurrences in window 17 [-239,-220] 27 -209.5 occurrences in window 18 [-219,-200] 28 -189.5 occurrences in window 19 [-199,-180] 29 -169.5 occurrences in window 20 [-179,-160] 30 -149.5 occurrences in window 21 [-159,-140] 31 -129.5 occurrences in window 22 [-139,-120] 32 -109.5 occurrences in window 23 [-119,-100] 33 -89.5 occurrences in window 24 [-99,-80] 34 -69.5 occurrences in window 25 [-79,-60] 35 -49.5 occurrences in window 26 [-59,-40] 36 -29.5 occurrences in window 27 [-39,-20] 37 -9.5 occurrences in window 28 [-19,0] 38 10.5 occurrences in window 29 [1,20] 39 30.5 occurrences in window 30 [21,40] 40 50.5 occurrences in window 31 [41,60] 41 70.5 occurrences in window 32 [61,80] 42 90.5 occurrences in window 33 [81,100] 43 110.5 occurrences in window 34 [101,120] 44 130.5 occurrences in window 35 [121,140] 45 150.5 occurrences in window 36 [141,160] 46 170.5 occurrences in window 37 [161,180] 47 190.5 occurrences in window 38 [181,200] 48 210.5 occurrences in window 39 [201,220] 49 230.5 occurrences in window 40 [221,240] 50 250.5 occurrences in window 41 [241,260] 51 270.5 occurrences in window 42 [261,280] 52 290.5 occurrences in window 43 [281,300] 53 310.5 occurrences in window 44 [301,320] 54 330.5 occurrences in window 45 [321,340] 55 350.5 occurrences in window 46 [341,360] 56 370.5 occurrences in window 47 [361,380] 57 390.5 occurrences in window 48 [381,400] 58 410.5 occurrences in window 49 [401,420] 59 430.5 occurrences in window 50 [421,440] 60 450.5 occurrences in window 51 [441,460] 61 470.5 occurrences in window 52 [461,480] 62 490.5 occurrences in window 53 [481,500] 63 510.5 occurrences in window 54 [501,520] 64 530.5 occurrences in window 55 [521,540] 65 550.5 occurrences in window 56 [541,560]
| seq | id | occ | chi2 | df | Pval | Eval | sig | rank | -549.5 | -529.5 | -509.5 | -489.5 | -469.5 | -449.5 | -429.5 | -409.5 | -389.5 | -369.5 | -349.5 | -329.5 | -309.5 | -289.5 | -269.5 | -249.5 | -229.5 | -209.5 | -189.5 | -169.5 | -149.5 | -129.5 | -109.5 | -89.5 | -69.5 | -49.5 | -29.5 | -9.5 | 10.5 | 30.5 | 50.5 | 70.5 | 90.5 | 110.5 | 130.5 | 150.5 | 170.5 | 190.5 | 210.5 | 230.5 | 250.5 | 270.5 | 290.5 | 310.5 | 330.5 | 350.5 | 370.5 | 390.5 | 410.5 | 430.5 | 450.5 | 470.5 | 490.5 | 510.5 | 530.5 | 550.5 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| aa | aa | 2052 | 108.2 | 55 | 2.5e-05 | 0.0004 | 3.40 | 1 | 1 | 8 | 2 | 7 | 6 | 14 | 33 | 19 | 22 | 36 | 32 | 39 | 39 | 53 | 31 | 46 | 50 | 48 | 32 | 44 | 43 | 33 | 43 | 48 | 73 | 66 | 68 | 86 | 63 | 83 | 77 | 79 | 69 | 41 | 55 | 37 | 45 | 51 | 45 | 44 | 40 | 45 | 34 | 27 | 48 | 33 | 24 | 19 | 20 | 18 | 21 | 1 | 5 | 1 | 5 | 0 |
| tt | tt | 2096 | 92.2 | 55 | 1.2e-03 | 0.02 | 1.71 | 2 | 0 | 4 | 2 | 8 | 8 | 14 | 8 | 18 | 24 | 22 | 31 | 18 | 22 | 17 | 33 | 39 | 36 | 52 | 49 | 47 | 62 | 57 | 67 | 78 | 81 | 69 | 57 | 73 | 66 | 59 | 49 | 84 | 71 | 71 | 67 | 70 | 45 | 56 | 66 | 52 | 24 | 30 | 34 | 44 | 29 | 28 | 23 | 24 | 30 | 26 | 17 | 16 | 8 | 9 | 2 | 0 |
| cc | cc | 808 | 82.4 | 55 | 9.8e-03 | 0.16 | 0.80 | 3 | 0 | 2 | 0 | 0 | 2 | 16 | 15 | 11 | 12 | 9 | 18 | 18 | 14 | 14 | 13 | 14 | 9 | 23 | 17 | 20 | 28 | 22 | 22 | 27 | 22 | 20 | 19 | 21 | 24 | 25 | 43 | 27 | 30 | 29 | 15 | 12 | 15 | 14 | 23 | 18 | 17 | 11 | 12 | 12 | 20 | 5 | 16 | 6 | 2 | 9 | 5 | 7 | 1 | 1 | 1 | 0 |
| ta | ta | 1398 | 71.1 | 55 | 7.1e-02 | 1.1 | -0.05 | 4 | 2 | 2 | 9 | 7 | 4 | 11 | 9 | 13 | 13 | 7 | 12 | 14 | 11 | 21 | 28 | 32 | 19 | 28 | 44 | 33 | 34 | 32 | 32 | 38 | 46 | 61 | 50 | 66 | 53 | 48 | 54 | 58 | 49 | 41 | 39 | 39 | 31 | 36 | 32 | 29 | 20 | 12 | 31 | 23 | 24 | 19 | 10 | 13 | 23 | 9 | 7 | 7 | 7 | 3 | 3 | 0 |
| gg | gg | 693 | 70.9 | 55 | 7.3e-02 | 1.2 | -0.07 | 5 | 0 | 1 | 0 | 0 | 6 | 4 | 13 | 9 | 12 | 11 | 7 | 20 | 11 | 6 | 11 | 4 | 22 | 12 | 9 | 19 | 7 | 21 | 31 | 15 | 26 | 22 | 25 | 20 | 29 | 23 | 28 | 19 | 13 | 20 | 20 | 21 | 18 | 16 | 17 | 13 | 10 | 13 | 13 | 6 | 7 | 15 | 12 | 8 | 6 | 7 | 9 | 3 | 1 | 1 | 0 | 1 |
| cg | cg | 751 | 56.7 | 55 | 4.1e-01 | 6.6 | -0.82 | 6 | 0 | 3 | 2 | 1 | 9 | 8 | 12 | 6 | 7 | 6 | 9 | 12 | 23 | 11 | 14 | 14 | 11 | 18 | 18 | 22 | 10 | 31 | 23 | 26 | 16 | 32 | 29 | 27 | 17 | 25 | 30 | 24 | 18 | 27 | 30 | 13 | 15 | 18 | 16 | 11 | 12 | 7 | 11 | 9 | 7 | 9 | 7 | 10 | 5 | 7 | 8 | 8 | 1 | 4 | 2 | 0 |
| ga | ga | 942 | 55.9 | 55 | 4.4e-01 | 7 | -0.85 | 7 | 0 | 1 | 2 | 5 | 7 | 8 | 12 | 11 | 10 | 14 | 12 | 14 | 19 | 11 | 18 | 19 | 25 | 29 | 33 | 19 | 23 | 30 | 27 | 30 | 33 | 25 | 42 | 33 | 41 | 40 | 33 | 29 | 26 | 20 | 24 | 31 | 25 | 10 | 15 | 10 | 13 | 13 | 9 | 13 | 12 | 18 | 11 | 11 | 2 | 7 | 5 | 5 | 4 | 1 | 1 | 1 |
| at | at | 1634 | 55.0 | 55 | 4.7e-01 | 7.6 | -0.88 | 8 | 0 | 2 | 6 | 10 | 8 | 11 | 17 | 17 | 18 | 16 | 14 | 14 | 20 | 23 | 33 | 33 | 30 | 39 | 40 | 32 | 48 | 37 | 37 | 46 | 52 | 68 | 64 | 77 | 70 | 51 | 65 | 55 | 64 | 42 | 37 | 43 | 36 | 36 | 41 | 37 | 21 | 18 | 28 | 25 | 26 | 20 | 13 | 17 | 23 | 15 | 15 | 10 | 7 | 5 | 2 | 0 |
| ca | ca | 1223 | 53.0 | 55 | 5.5e-01 | 8.8 | -0.95 | 9 | 0 | 3 | 3 | 5 | 9 | 8 | 14 | 18 | 12 | 14 | 26 | 24 | 15 | 21 | 20 | 16 | 32 | 22 | 24 | 25 | 38 | 37 | 29 | 31 | 41 | 55 | 49 | 30 | 48 | 46 | 41 | 30 | 38 | 30 | 25 | 21 | 32 | 36 | 29 | 21 | 25 | 30 | 20 | 18 | 26 | 12 | 15 | 11 | 12 | 15 | 7 | 7 | 3 | 2 | 2 | 0 |
| ag | ag | 991 | 51.7 | 55 | 6.0e-01 | 9.6 | -0.98 | 10 | 1 | 2 | 3 | 5 | 5 | 12 | 7 | 10 | 8 | 11 | 20 | 18 | 14 | 12 | 21 | 20 | 22 | 29 | 31 | 26 | 15 | 35 | 26 | 29 | 37 | 29 | 37 | 26 | 34 | 47 | 25 | 35 | 27 | 23 | 24 | 29 | 25 | 19 | 17 | 14 | 21 | 20 | 15 | 15 | 17 | 17 | 16 | 10 | 6 | 10 | 1 | 4 | 6 | 1 | 2 | 0 |
| ct | ct | 1004 | 50.5 | 55 | 6.5e-01 | 10 | -1.02 | 11 | 0 | 2 | 2 | 1 | 7 | 5 | 5 | 10 | 9 | 13 | 13 | 15 | 11 | 17 | 8 | 14 | 16 | 16 | 30 | 33 | 27 | 15 | 33 | 40 | 37 | 30 | 31 | 34 | 32 | 37 | 40 | 41 | 29 | 31 | 25 | 36 | 23 | 22 | 17 | 21 | 25 | 22 | 17 | 21 | 13 | 15 | 9 | 13 | 10 | 15 | 5 | 3 | 1 | 1 | 6 | 0 |
| gc | gc | 987 | 50.3 | 55 | 6.6e-01 | 10 | -1.02 | 12 | 0 | 3 | 3 | 2 | 11 | 11 | 10 | 11 | 12 | 13 | 19 | 22 | 14 | 14 | 21 | 19 | 23 | 17 | 21 | 30 | 24 | 35 | 23 | 30 | 27 | 39 | 40 | 22 | 20 | 40 | 26 | 30 | 27 | 29 | 25 | 23 | 27 | 22 | 24 | 14 | 22 | 14 | 17 | 12 | 15 | 11 | 15 | 8 | 8 | 17 | 8 | 7 | 4 | 3 | 3 | 0 |
| tg | tg | 1229 | 46.8 | 55 | 7.8e-01 | 12 | -1.09 | 13 | 0 | 1 | 0 | 5 | 8 | 8 | 16 | 15 | 16 | 18 | 16 | 20 | 17 | 13 | 15 | 19 | 26 | 25 | 22 | 35 | 45 | 31 | 44 | 49 | 42 | 34 | 48 | 43 | 47 | 34 | 37 | 35 | 40 | 36 | 27 | 38 | 41 | 26 | 26 | 21 | 16 | 12 | 12 | 21 | 17 | 19 | 15 | 12 | 16 | 17 | 13 | 10 | 6 | 1 | 2 | 1 |
| gt | gt | 1057 | 46.3 | 55 | 7.9e-01 | 13 | -1.10 | 14 | 2 | 1 | 2 | 6 | 6 | 11 | 11 | 8 | 9 | 11 | 16 | 15 | 18 | 15 | 13 | 12 | 16 | 27 | 20 | 32 | 25 | 32 | 46 | 44 | 41 | 30 | 31 | 40 | 36 | 30 | 32 | 30 | 32 | 36 | 30 | 25 | 32 | 28 | 19 | 24 | 10 | 12 | 14 | 18 | 10 | 18 | 13 | 13 | 16 | 10 | 10 | 9 | 6 | 2 | 2 | 0 |
| tc | tc | 1062 | 45.5 | 55 | 8.2e-01 | 13 | -1.12 | 15 | 0 | 2 | 1 | 4 | 8 | 8 | 6 | 9 | 9 | 10 | 17 | 13 | 19 | 19 | 11 | 9 | 16 | 26 | 28 | 25 | 20 | 23 | 35 | 44 | 42 | 36 | 32 | 36 | 40 | 33 | 45 | 36 | 35 | 32 | 29 | 23 | 23 | 23 | 20 | 28 | 27 | 25 | 15 | 21 | 11 | 14 | 9 | 16 | 14 | 12 | 7 | 7 | 0 | 4 | 5 | 0 |
| ac | ac | 935 | 39.9 | 55 | 9.4e-01 | 15 | -1.18 | 16 | 0 | 3 | 3 | 2 | 6 | 4 | 12 | 15 | 7 | 9 | 12 | 19 | 13 | 16 | 10 | 18 | 18 | 17 | 24 | 22 | 33 | 29 | 21 | 25 | 27 | 44 | 38 | 26 | 40 | 39 | 35 | 28 | 26 | 24 | 28 | 20 | 28 | 27 | 17 | 11 | 16 | 16 | 18 | 15 | 18 | 13 | 8 | 9 | 7 | 6 | 6 | 4 | 1 | 1 | 1 | 0 |
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