; matrix-scan -v 1 -matrix_format transfac -m $RSAT/public_html/tmp/www-data/2026/06/17/matrix-scan_2026-06-17.061147_81damk.matrix -pseudo 1 -decimals 1 -2str -origin end -offset -200 -bginput -markov 1 -bg_pseudo 0.01 -return limits -return sites -return pval -return rank -lth score 1 -uth pval 1e-4 -i $RSAT/public_html/tmp/www-data/2026/06/17/tmp_sequence_2026-06-17.061147_Fz4NoV.fasta -seq_format fasta -n score ; Slow counting mode ; Input files ; input $RSAT/public_html/tmp/www-data/2026/06/17/tmp_sequence_2026-06-17.061147_Fz4NoV.fasta ; Matrix files ; matrix 1 $RSAT/public_html/tmp/www-data/2026/06/17/matrix-scan_2026-06-17.061147_81damk.matrix ; Sequence format fasta ; Pseudo counts 1 ; Background model ; Method input ; Markov order 1 ; Strand undef ; Background pseudo-frequency 0.01 ; Residue probabilities ; a 0.34242 ; c 0.18661 ; g 0.15145 ; t 0.31953 ; Thresholds lower upper ; pval NA 0.0001 ; score 1 NA ; Output columns ; 1 seq_id ; 2 ft_type ; 3 ft_name ; 4 strand ; 5 start ; 6 end ; 7 sequence ; 8 weight #seq_id ft_type ft_name strand start end sequence weight Pval ln_Pval sig rank rank_pm BVRB_3g059400|Beta_vulgaris.RefBeet-1.2.2.60|BVRB_3g059400 limit START_END D -255 199 . 0 0 0 0 BVRB_3g059400|Beta_vulgaris.RefBeet-1.2.2.60|BVRB_3g059400 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -137 -119 GTTCATGAATCATGCCTTT 10.0 4.9e-06 -12.228 5.311 1 1 BVRB_4g096340|Beta_vulgaris.RefBeet-1.2.2.60|BVRB_4g096340 limit START_END D -1501 199 . 0 0 0 0 BVRB_4g096340|Beta_vulgaris.RefBeet-1.2.2.60|BVRB_4g096340 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -1330 -1312 NNNNNNNGGTCGGACCGTG 7.5 3.8e-05 -10.175 4.419 1 1 T459_03445|Capsicum_annuum.ASM51225v2.60|T459_03445 limit START_END D -1501 199 . 0 0 0 0 T459_31204|Capsicum_annuum.ASM51225v2.60|T459_31204 limit START_END D -1501 199 . 0 0 0 0 T459_31204|Capsicum_annuum.ASM51225v2.60|T459_31204 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -844 -826 TAGCATGGACAATGCCATC 7.7 3.3e-05 -10.327 4.485 1 1 T459_31204|Capsicum_annuum.ASM51225v2.60|T459_31204 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -128 -110 CTGAATGTTTCATGCCACT 6.8 6.4e-05 -9.657 4.194 2 2 AUR62023693.v1.0|Chenopodium_quinoa.PI614886.392.v1.JGI|AUR62023693.v1.0 limit START_END D -1501 199 . 0 0 0 0 AUR62023693.v1.0|Chenopodium_quinoa.PI614886.392.v1.JGI|AUR62023693.v1.0 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -70 -52 CGGCATGCTTCGAGCCTTG 16.3 2.7e-09 -19.718 8.564 1 1 AUR62023693.v1.0|Chenopodium_quinoa.PI614886.392.v1.JGI|AUR62023693.v1.0 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -68 -50 AGGCTCGAAGCATGCCGCT 9.1 1.1e-05 -11.446 4.971 2 2 AUR62023693.v1.0|Chenopodium_quinoa.PI614886.392.v1.JGI|AUR62023693.v1.0 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -106 -88 GCGGATGATTCATGCCTTT 7.8 3.0e-05 -10.404 4.518 3 3 Cla97C08G153070|Citrullus_lanatus.Cla97_v1.62|Cla97C08G153070 limit START_END D -1501 199 . 0 0 0 0 Cla97C08G153070|Citrullus_lanatus.Cla97_v1.62|Cla97C08G153070 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -866 -848 CATCATGGAGATCGCCAAA 9.1 1.1e-05 -11.446 4.971 1 1 MELO3C011439.2|Cucumis_melo.Melonv4.60|MELO3C011439.2 limit START_END D -1501 199 . 0 0 0 0 MELO3C011439.2|Cucumis_melo.Melonv4.60|MELO3C011439.2 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -693 -675 AAGCACGAATCAAGCCATG 8.8 1.4e-05 -11.197 4.863 1 1 MELO3C011439.2|Cucumis_melo.Melonv4.60|MELO3C011439.2 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -691 -673 TGGCTTGATTCGTGCTTCT 6.9 5.9e-05 -9.737 4.229 2 2 Csa_6G366300|Cucumis_sativus.ASM407v2.60|Csa_6G366300 limit START_END D -1501 199 . 0 0 0 0 HanXRQr2_Chr13g0574171|Helianthus_annuus.HanXRQr2.0-SUNRISE.60|HanXRQr2_Chr13g0574171 limit START_END D -1501 199 . 0 0 0 0 HanXRQr2_Chr13g0574171|Helianthus_annuus.HanXRQr2.0-SUNRISE.60|HanXRQr2_Chr13g0574171 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -175 -157 ATGCATGAGAATGGCCTTG 11.7 9.9e-07 -13.821 6.002 1 1 HanXRQr2_Chr13g0574171|Helianthus_annuus.HanXRQr2.0-SUNRISE.60|HanXRQr2_Chr13g0574171 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -74 -56 ATGCATGAGAATGGCCTTA 11.0 2.0e-06 -13.146 5.709 2 2 HanXRQr2_Chr13g0574181|Helianthus_annuus.HanXRQr2.0-SUNRISE.60|HanXRQr2_Chr13g0574181 limit START_END D -1501 199 . 0 0 0 0 gene-LSAT_8X33320|Lactuca_sativa.Lsat_Salinas_v7.60|gene-LSAT_8X33320 limit START_END D -1501 199 . 0 0 0 0 gene-LSAT_8X33320|Lactuca_sativa.Lsat_Salinas_v7.60|gene-LSAT_8X33320 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -207 -189 TTGCATGAAACATGCCAAT 10.9 2.1e-06 -13.052 5.668 1 1 gene-LSAT_8X33320|Lactuca_sativa.Lsat_Salinas_v7.60|gene-LSAT_8X33320 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -154 -136 TGTCATGTTACATGGCTTT 6.6 7.3e-05 -9.520 4.135 2 2 gene25542|Medicago_truncatula.MtrunA17r50ANR.60|gene25542 limit START_END D -1501 199 . 0 0 0 0 107761937|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107761937 limit START_END D -1501 199 . 0 0 0 0 107761937|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107761937 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -130 -112 AGGCATGAGACAGGCCGAG 17.0 7.8e-10 -20.970 9.107 1 1 107761937|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107761937 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -132 -114 CGGCCTGTCTCATGCCTTT 12.9 2.8e-07 -15.104 6.560 2 2 107761937|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107761937 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -379 -361 CGTAATGGATCGGGCCAGA 10.8 2.4e-06 -12.948 5.623 3 3 107761937|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107761937 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -79 -61 GGGCTTGGAACGCCCGTTT 10.1 4.5e-06 -12.308 5.345 4 4 107761937|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107761937 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -428 -410 GATCTTGGCGGGTACCTCG 7.2 4.8e-05 -9.950 4.321 5 5 107793142|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107793142 limit START_END D -1501 199 . 0 0 0 0 107793142|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107793142 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D 30 48 CAGCATGGGGCAGGCCGAA 15.5 9.6e-09 -18.457 8.016 1 1 107793142|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107793142 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D 55 73 AGGCATAGGACATGCCAAG 12.6 3.9e-07 -14.762 6.411 2 2 107793142|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107793142 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R 53 71 TGGCATGTCCTATGCCTTT 9.8 5.9e-06 -12.044 5.230 3 3 107793142|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107793142 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R 28 46 CGGCCTGCCCCATGCTGGA 6.6 7.3e-05 -9.520 4.135 4 4 107759548|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107759548 limit START_END D -1501 199 . 0 0 0 0 107759548|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107759548 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -51 -33 AGGCATGGGACATGCCGAG 18.0 1.0e-10 -22.981 9.980 1 1 107759548|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107759548 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -76 -58 CGGCATGGGACAGGCCGAA 17.5 3.0e-10 -21.931 9.524 2 2 107759548|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107759548 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -53 -35 CGGCATGTCCCATGCCTTT 13.8 9.7e-08 -16.145 7.012 3 3 107759548|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107759548 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -78 -60 CGGCCTGTCCCATGCCGGA 10.8 2.4e-06 -12.948 5.623 4 4 107759548|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107759548 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -1311 -1293 CGGGATGGGACGGGACAAA 8.5 1.7e-05 -10.962 4.761 5 5 107759548|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107759548 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -1217 -1199 CGGAATGAAACAGGACGGG 7.6 3.5e-05 -10.258 4.455 6 6 107759548|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107759548 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -1168 -1150 CGGAATGGGACGGGACGGC 7.2 4.8e-05 -9.950 4.321 7 7 107759548|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107759548 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -1340 -1322 GGACGTGGGATAGGCGGGA 6.5 7.9e-05 -9.442 4.100 8 8 107759548|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107759548 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -1158 -1140 CGGGACGGCCCGTCCCGTT 6.5 7.9e-05 -9.442 4.100 9 9 107759548|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107759548 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -1328 -1310 CGTCATGTCCCGTCCCGCC 6.3 9.1e-05 -9.307 4.042 10 10 107812499|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107812499 limit START_END D -1501 199 . 0 0 0 0 107812499|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107812499 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -114 -96 AGGCATGGGACGGGCCGAG 19.1 5.5e-12 -25.919 11.256 1 1 107812499|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107812499 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -65 -47 GGGCTTGGAACGGCCGTTT 10.8 2.4e-06 -12.948 5.623 2 2 107812499|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107812499 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -116 -98 CGGCCCGTCCCATGCCTTT 8.2 2.2e-05 -10.715 4.654 3 3 gene-LOC110794951|Spinacia_oleracea.GCF020520425.1.RefSeq|gene-LOC110794951 limit START_END D -1501 199 . 0 0 0 0 gene-LOC110794951|Spinacia_oleracea.GCF020520425.1.RefSeq|gene-LOC110794951 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R 45 63 CGGCGTGCTTCGAGCTGTT 10.0 4.9e-06 -12.228 5.311 1 1 gene-LOC110794951|Spinacia_oleracea.GCF020520425.1.RefSeq|gene-LOC110794951 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R 113 131 CGGATTGGTAAGTGCGTGG 8.9 1.3e-05 -11.288 4.902 2 2 gene-LOC110794951|Spinacia_oleracea.GCF020520425.1.RefSeq|gene-LOC110794951 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D 47 65 CAGCTCGAAGCACGCCGCT 7.6 3.5e-05 -10.258 4.455 3 3 gene-LOC110794951|Spinacia_oleracea.GCF020520425.1.RefSeq|gene-LOC110794951 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D 3 21 AGGCATGAAACATCGCAAA 7.0 5.5e-05 -9.803 4.257 4 4 AT3G13610|Arabidopsis_thaliana.TAIR10.60|F6'H1 limit START_END D -1501 199 . 0 0 0 0 AT3G13610|Arabidopsis_thaliana.TAIR10.60|F6'H1 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D 24 42 AAGCATGGTGAGAGCCATA 13.3 1.7e-07 -15.561 6.758 1 1 AT3G13610|Arabidopsis_thaliana.TAIR10.60|F6'H1 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -1482 -1464 AAGCGTGAGACTAGGCATG 7.3 4.4e-05 -10.032 4.357 2 2 GSBRNA2T00023848001|Brassica_napus.AST_PRJEB5043_v1.60|BnaC05g39690D limit START_END D -1448 199 . 0 0 0 0 GSBRNA2T00023848001|Brassica_napus.AST_PRJEB5043_v1.60|BnaC05g39690D site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -891 -873 AGGCATGGACTATGCATGG 7.8 3.0e-05 -10.404 4.518 1 1 GSBRNA2T00023848001|Brassica_napus.AST_PRJEB5043_v1.60|BnaC05g39690D site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -889 -871 ATGCATAGTCCATGCCTCT 7.2 4.8e-05 -9.950 4.321 2 2 GSBRNA2T00104708001|Brassica_napus.AST_PRJEB5043_v1.60|BnaA05g25780D limit START_END D -1501 199 . 0 0 0 0 GSBRNA2T00104708001|Brassica_napus.AST_PRJEB5043_v1.60|BnaA05g25780D site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -899 -881 AGGCATGGACTATGCATGG 7.8 3.0e-05 -10.404 4.518 1 1 GSBRNA2T00104708001|Brassica_napus.AST_PRJEB5043_v1.60|BnaA05g25780D site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -897 -879 ATGCATAGTCCATGCCTCT 7.2 4.8e-05 -9.950 4.321 2 2 GSBRNA2T00123637001|Brassica_napus.AST_PRJEB5043_v1.60|BnaC03g38030D limit START_END D -1501 199 . 0 0 0 0 GSBRNA2T00123637001|Brassica_napus.AST_PRJEB5043_v1.60|BnaC03g38030D site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -134 -116 AAGCATGGTGAGAGCCATA 13.3 1.7e-07 -15.561 6.758 1 1 GSBRNA2T00123637001|Brassica_napus.AST_PRJEB5043_v1.60|BnaC03g38030D site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -1244 -1226 AAGCATGGGACTGGGCATG 9.9 5.3e-06 -12.140 5.272 2 2 GSBRNA2T00137685001|Brassica_napus.AST_PRJEB5043_v1.60|BnaA03g32900D limit START_END D -1459 199 . 0 0 0 0 GSBRNA2T00137685001|Brassica_napus.AST_PRJEB5043_v1.60|BnaA03g32900D site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -144 -126 AAGCATGGTGAGAGCCATA 13.3 1.7e-07 -15.561 6.758 1 1 GSBRNA2T00137685001|Brassica_napus.AST_PRJEB5043_v1.60|BnaA03g32900D site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -988 -970 AAGCATGGTACTGGGCATG 9.5 7.6e-06 -11.792 5.121 2 2 A03p040480.1_BraROA|Brassica_rapa_ro18.SCU_BraROA_2.3.60|A03p040480.1_BraROA limit START_END D -1426 199 . 0 0 0 0 A03p040480.1_BraROA|Brassica_rapa_ro18.SCU_BraROA_2.3.60|A03p040480.1_BraROA site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -144 -126 AAGCATGGTGAGAGCCATA 13.3 1.7e-07 -15.561 6.758 1 1 GLYMA_07G124400|Glycine_max.Glycine_max_v2.1.60|GLYMA_07G124400 limit START_END D -1501 199 . 0 0 0 0 gene-Lalb_Chr13g0290931|Lupinus_albus.CNRS_Lalb_10.GCA_009771035.1.NCBI|gene-Lalb_Chr13g0290931 limit START_END D -1501 199 . 0 0 0 0 gene-Lalb_Chr13g0290931|Lupinus_albus.CNRS_Lalb_10.GCA_009771035.1.NCBI|gene-Lalb_Chr13g0290931 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -171 -153 AAACATGTAACTTGCCATT 8.2 2.2e-05 -10.715 4.654 1 1 gene-Lalb_Chr13g0290931|Lupinus_albus.CNRS_Lalb_10.GCA_009771035.1.NCBI|gene-Lalb_Chr13g0290931 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -1037 -1019 CTGCATGAGTAANNNNNNN 6.1 1.0e-04 -9.167 3.981 2 2 107763182|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107763182 limit START_END D -1501 199 . 0 0 0 0 107763182|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107763182 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -893 -875 CTGAATGGCCAGTGCCAAA 8.6 1.6e-05 -11.035 4.792 1 1 107785648|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107785648 limit START_END D -1501 199 . 0 0 0 0 107796566|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107796566 limit START_END D -1501 199 . 0 0 0 0 107796566|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107796566 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 D -42 -24 TGGCATGCCAATTGCCTAT 11.4 1.3e-06 -13.533 5.877 1 1 107796566|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107796566 site dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 R -44 -26 AGGCAATTGGCATGCCATG 6.3 9.1e-05 -9.307 4.042 2 2 107787976|Nicotiana_tabacum_GCF_000715075.1_ASM71507v2|LOC107787976 limit START_END D -1501 199 . 0 0 0 0 Prudul26B000915|Prunus_dulcis.ALMONDv2.60|Prudul26B000915 limit START_END D -773 199 . 0 0 0 0 PRUPE_6G006000|Prunus_persica.Prunus_persica_NCBIv2.60|PRUPE_6G006000 limit START_END D -233 199 . 0 0 0 0 Solyc11g072100.ITAG4.0|Solanum_lycopersicum.ITAG4.0.JGI|Solyc11g072100.ITAG4.0 limit START_END D -1501 199 . 0 0 0 0 ; ; Matrices ; matrix name ncol nrow pseudo Wmin Wmax Wrange ; 1 dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 19 4 1 -37.100 20.000 57.100 a:0.342 c:0.187 g:0.151 t:0.320 ; Number of sequences scanned 32 ; Sum of sequence lengths 51020 ; N residues 10767 ; Matches per matrix ; matrix name matches scored ; 1 dyads_test_vs_ctrl_m1dyads_test_vs_ctrl_m1 56 100888 ; TOTAL 56 100888 ; Host name rsat ; Job started 2026-06-17.061147 ; Job done 2026-06-17.061159 ; Seconds 11.74 ; user 11.74 ; system 0.01 ; cuser 0.12 ; csystem 0.01